Draba brachycarpa |
Draba cinerea |
|
---|---|---|
short-pod whitlowgrass, shortfruit Draba, shortpod Draba |
gray-leaf Draba, gray-leaf whitlow-grass |
|
Habit | Annuals; not scapose. | Perennials; (cespitose); caudex simple or branched; rarely scapose. |
Stems | usually branched, rarely unbranched, (0.3–)0.4–1.9(–2.2) dm, pubescent throughout, trichomes sessile, cruciform, 0.2–0.6(–0.8) mm (rays often equal, or those parallel to stem axis longer). |
unbranched, (0.3–)0.5–1.6(–2.7) dm, pubescent throughout, trichomes 4–10-rayed, 0.1–0.3 mm, (often some rays branched, simple trichomes sparse, to 0.6 mm). |
Basal leaves | not rosulate; petiolate; petiole (to 0.5 cm), not ciliate; blade oblanceolate to obovate, 0.5–2 cm × 2–8 mm, margins entire, surfaces pubescent, trichomes cruciform, 0.2–0.6(–0.8) mm. |
rosulate; petiolate; petiole base and proximalmost margin ciliate, (trichomes simple, 0.2–0.8 mm); blade oblanceolate to narrowly obovate or linear-lanceolate, 0.4–1.5 cm × 1–5 mm, margins usually entire, rarely with 1 tooth on each side, surfaces densely pubescent with minutely stalked, stellate, 8–12-rayed, trichomes 0.1–0.25 mm, (midvein obscure abaxially, apex trichomes simple). |
Cauline leaves | (4–)6–11; sessile; blade lanceolate to oblong or linear, margins entire, surfaces pubescent as basal. |
0–3(–5); sessile; blade ovate or oblong to lanceolate, margins entire, surfaces pubescent as basal. |
Racemes | (main branch) (20–)25–65(–74)-flowered, ebracteate, elongated in fruit; rachis not flexuous, pubescent as stem. |
(3–)5–17(–24)-flowered, ebracteate, elongated in fruit; rachis not flexuous, pubescent as stem. |
Flowers | (late ones cleistogamous, apetalous); sepals (green or pink), oblong, 0.7–1.2(–1.5) mm, pubescent; petals white, spatulate, 2–3 × 0.8–1.1 mm; anthers ovate, 0.15–0.25 mm. |
sepals ovate, 1.7–2.5 mm, pubescent, (trichomes simple and short-stalked, 2- or 3-rayed); petals white, spatulate to obovate, 3.5–4.5 × 1.5–2 mm; anthers ovate, 0.3–0.4 mm. |
Fruiting pedicels | horizontal to divaricate-ascending, straight, (1–)1.5–4(–5) mm, pubescent, trichomes cruciform. |
divaricate-ascending or ascending, straight, (3–)4–7(–9) mm, pubescent as stem. |
Fruits | usually elliptic to linear-elliptic, rarely ovate-elliptic, plane, flattened, (2–)2.5–5(–6) × 0.9–1.4(–1.9) mm; valves glabrous; ovules 8–16(–20) per ovary; style 0.05–0.1 mm. |
oblong to elliptic, plane, slightly flattened, 5–8 × 2–3 mm; valves pubescent, trichomes short-stalked, 2–5-rayed, 0.05–0.3 mm, (some rays branched); ovules 20–36(–42) per ovary; style (0.1–)0.2–1 mm. |
Seeds | (winged), ovoid, 0.5–0.7 × 0.4–0.5 mm. |
ovoid, 0.6–0.8 × 0.4–0.6 mm. |
2n | = 16. |
= 48. |
Draba brachycarpa |
Draba cinerea |
|
Phenology | Flowering Feb–May. | Flowering Jun–Jul. |
Habitat | Open woods, cedar glades, pastures and lawns, roadsides, disturbed sites | Rock outcrops, ridges, meadows, gravel beaches, stream banks, alluvial fans |
Elevation | 0-300 m (0-1000 ft) | 0-500 m (0-1600 ft) |
Distribution |
AL; AR; AZ; FL; GA; IL; IN; KS; KY; LA; MO; MS; NC; OH; OK; OR; SC; TN; TX; VA
|
AK; BC; NT; NU; ON; QC; SK; YT; Greenland; Europe (Finland, Norway, n Russia); e Asia (Russian Far East, Siberia) |
Discussion | Draba brachycarpa is closely related to D. aprica and the two are sometimes confused. It is readily distinguished by having leaves with sessile (versus stalked) trichomes, glabrous (versus pubescent) fruits, and smaller (0.5–0.7 versus 0.9–1.1 mm) seeds. The records from Arizona and Oregon are based on old collections, Porter 802 (Devil’s Canyon, 22 Feb 1926, US) and Howell s.n. (near Cobarg, Willamette Valley, 7 Apr 1887, US); it is not known if these records represent introductions or remnants of a previously wider distribution. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Draba cinerea is a polymorphic species in which tetraploid, hexaploid, heptaploid, and octoploid chromosome counts (all based on x = 8) have been reported (e.g., R. C. Rollins 1993). Although we have not seen the type collection, we tentatively consider the taxon to be hexaploid, which is the predominant ploidy level throughout the species range, including North America, and also from its type region in Siberia. Hexaploid Draba cinerea is closely related to decaploid (2n = 80) D. arctica, from which it is distinguished with difficulty. In general, D. cinerea has 1–3(–5)-leaved stems, basal leaf blades usually not terminated with simple trichomes and with obscure midveins abaxially, petals 3.5–4.5 mm, and seeds 0.6–0.8 × 0.4–0.6 mm. By contrast, D. arctica has leafless or 1(–3)-leaved stems, basal leaf blades terminated with simple or 2-rayed trichomes and with midveins distinct abaxially, petals 3.5–6 mm, and seeds (0.8–)0.9–1.1 × (0.6–)0.7–0.8 mm. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 296. | FNA vol. 7, p. 300. |
Parent taxa | Brassicaceae > tribe Arabideae > Draba | Brassicaceae > tribe Arabideae > Draba |
Sibling taxa | ||
Synonyms | Abdra brachycarpa | D. magellanica subsp. cinerea |
Name authority | Nuttall ex Torrey & A. Gray: Fl. N. Amer. 1: 108. (1838) | Adams: Mém Soc. Imp. Naturalistes Moscou 5: 103. (1817) |
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