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golden Draba, golden Draba whitlow-grass, golden whitlow-grass, twisted-pod Draba

alpine tundra Draba

Habit Perennials; (not cespitose); caudex simple or branched; not scapose. Perennials; caudex branched (often with persistent leaf remains, branches not creeping); not scapose.
Stems

sometimes branched distally, (0.5–) 1–3.5(–5.2) dm, pubescent throughout, trichomes simple (non-crisped), 0.4–1.3 mm, and 3–6-rayed ones, 0.1–0.5 mm.

unbranched, (0.1–)0.2–1(–1.3) dm, pubescent throughout, trichomes subsessile (often crisped), 3–5-rayed, stellate, 0.03–0.25(–0.4) mm, (rays sometimes forked).

Basal leaves

rosulate; petiolate;

petiole (distinct or obscure) ciliate, (trichomes simple, to 0.8 mm);

blade oblanceolate to obovate, (0.4–)1–3.7(–5) cm × (1–)2–7(–10) mm, margins entire or denticulate, surfaces pubescent, trichomes stalked, (2–)4–7 (or 8)-rayed, 0.2–0.5(–0.6) mm.

rosulate;

petiole (obscure), usually not ciliate, rarely sparsely pubescent, (trichomes simple, to 0.6 mm);

blade oblanceolate to linear-oblanceolate, (0.4–)0.5–3(–4) cm × 1–5 mm, margins entire, surfaces pubescent with short-stalked (crisped), 3–8-rayed trichomes, 0.05–0.4 mm.

Cauline leaves

5–20(–26);

sessile;

blade oblong to lanceolate or ovate, margins entire or dentate, surfaces pubescent as basal, sometimes adaxially with simple trichomes.

(1 or) 2–4 (or 5);

sessile;

blade oblong to ovate or linear, margins entire, surfaces pubescent as basal.

Racemes

(10–)18–52(–72)-flowered, usually bracteate on proximalmost 1–12(–17) flowers, rarely ebracteate, elongated in fruit;

rachis not flexuous, pubescent as stem.

4–10(–18)-flowered, ebracteate, elongated in fruit;

rachis not flexuous, pubescent as stem.

Flowers

sepals (green or yellowish), oblong, 2.2–3 mm, pubescent, (trichomes simple and branched);

petals yellow, oblanceolate, 3.5–5 × 1.5–2.5 mm;

anthers ovate, 0.4–0.5 mm.

sepals ovate, 2–3 mm, pubescent, (trichomes simple and short-stalked, 2–4-rayed);

petals yellow, spatulate, 3–5 × 1.5–3 mm, (clawed);

anthers ovate, 0.25–0.4 mm.

Fruiting pedicels

divaricate-ascending to ascending or suberect, straight, 3–13(–20) mm, pubescent as stem.

ascending, usually straight, rarely curved upward, (2–)3–8(–12) mm, pubescent as stem.

Fruits

(often subappressed to rachis), lanceolate to linear-lanceolate or narrowly oblong, slightly twisted or plane, flattened, (6–)9–14(–17) × 2–3.5 mm;

valves pubescent, trichomes simple and short-stalked, 2–4-rayed, 0.05–0.3 mm;

ovules 28–38(–44) per ovary;

style 0.5–1.2(–1.5) mm.

(not appressed to rachis), ovate to elliptic or lanceolate, slightly twisted or plane, flattened, (3–)5–10 × 2–4 mm;

valves often pubescent, occasionally glabrous, trichomes simple and minutely stalked, 2–4-rayed, 0.03–0.25 mm;

ovules 10–16(–18) per ovary;

style 0.3–0.8(–1.2) mm.

Seeds

oblong, 0.9–1.3 × 0.5–0.7 mm.

oblong, 1–1.6 × 0.6–1 mm.

2n

= 74.

= ca. 64.

Draba aurea

Draba streptobrachia

Phenology Flowering Jun–Aug. Flowering Jul–Aug.
Habitat Rock outcrops, talus, damp gullies and meadows, subalpine conifer woodlands, alpine slopes and turf, tundra, road banks, river gravel Alpine tundra, scree, ridges and alpine slopes, turf, fellfields, talus slopes, crevices in rock ledges, loose soils
Elevation (0-)700-4200 m ((0-)2300-13800 ft) 3200-4000 m (10500-13100 ft)
Distribution
from FNA
AK; AZ; CO; ID; MT; NM; NV; SD; UT; WA; WY; AB; BC; MB; NF; NT; NU; ON; QC; SK; YT; Greenland
[WildflowerSearch map]
[BONAP county map]
from FNA
CO
[WildflowerSearch map]
[BONAP county map]
Discussion

Draba aurea is extremely variable in plant size, number of cauline leaves, number of bracteate flowers, style length, and fruit size, shape, orientation, twisting, and indumentum. Much of the variation in the number of bracts, style length, fruit twisting, and growth habit occurs in Greenland, where the type specimen was collected and where the species is found near sea level.

The highly deviant chromosome counts (e.g., 2n = 40 + 1, 64, 82) listed by R. C. Rollins (1993) and S. I. Warwick and I. A. Al-Shehbaz (2006) are mostly unvouchered and have to be disregarded; counts of 2n = ca. 80 have been re-assigned to Draba glabella. Published (G. A. Mulligan 2002) and unpublished counts made by Mulligan and M. D. Windham from Alaska, British Columbia, Colorado, Quebec, Utah, and Yukon indicate that the most common chromosome number of D. aurea is 2n = 74 (or 72). This suggests that the species is an allopolyploid (hexaploid or higher), incorporating genomes from both euploid and aneuploid lineages (M. A. Beilstein and Windham 2003). Detailed cytological and molecular studies are much needed to fully understand this widely distributed and highly variable species.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

As indicated by Price, Draba streptobrachia is an apomict, yielding abundant, well-developed seed despite producing only abortive pollen. Morphological studies (M. D. Windham, unpubl.) suggest that the species may be an allopolyploid containing a genome from D. crassa.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 294. FNA vol. 7, p. 341.
Parent taxa Brassicaceae > tribe Arabideae > Draba Brassicaceae > tribe Arabideae > Draba
Sibling taxa
D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. cyclomorpha, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. helleriana, D. hitchcockii, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis
D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. cyclomorpha, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. helleriana, D. hitchcockii, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis
Synonyms D. aurea var. aureiformis, D. aurea var. leiocarpa, D. aurea var. luteola, D. aureiformis, D. aureiformis var. leiocarpa, D. bakeri, D. decumbens, D. henneana var. maccallae, D. luteola, D. luteola var. minganensis, D. maccallae, D. minganensis, D. surculifera, D. uber D. chrysantha, D. spectabilis var. dasycarpa
Name authority Vahl ex Hornemann: Fors. Oecon. Plantel. ed. 2, 599. (1806) R. A. Price: Brittonia 32: 168. (1980)
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