Draba aurea |
Draba sibirica |
|
---|---|---|
golden Draba, golden Draba whitlow-grass, golden whitlow-grass, twisted-pod Draba |
Siberian whitlowgrass |
|
Habit | Perennials; (not cespitose); caudex simple or branched; not scapose. | Perennials; (stoloniferous); caudex branched (sparsely covered with persistent petiole remains, branches slender, creeping); scapose. |
Stems | sometimes branched distally, (0.5–) 1–3.5(–5.2) dm, pubescent throughout, trichomes simple (non-crisped), 0.4–1.3 mm, and 3–6-rayed ones, 0.1–0.5 mm. |
(subdecumbent when sterile), unbranched, 0.5–2.5 dm, sparsely to densely hispid, trichomes malpighiaceous, (flowering scapes sparsely pilose proximally with simple trichomes, often glabrous on distal 1/2). |
Basal leaves | rosulate; petiolate; petiole (distinct or obscure) ciliate, (trichomes simple, to 0.8 mm); blade oblanceolate to obovate, (0.4–)1–3.7(–5) cm × (1–)2–7(–10) mm, margins entire or denticulate, surfaces pubescent, trichomes stalked, (2–)4–7 (or 8)-rayed, 0.2–0.5(–0.6) mm. |
rosulate; petiolate; blade oblong-lanceolate or oblanceolate, 0.4–2.2 cm × 1–5(–10) mm, margins entire, surfaces often pubescent, sometimes glabrous (except margins), with appressed, malpighiaceous trichomes (these sometimes with 1 or 2 shorter, lateral branches, appearing 3-fid or cross-shaped). |
Cauline leaves | 5–20(–26); sessile; blade oblong to lanceolate or ovate, margins entire or dentate, surfaces pubescent as basal, sometimes adaxially with simple trichomes. |
0. |
Racemes | (10–)18–52(–72)-flowered, usually bracteate on proximalmost 1–12(–17) flowers, rarely ebracteate, elongated in fruit; rachis not flexuous, pubescent as stem. |
7–20-flowered, ebracteate, considerably elongated in fruit; rachis not flexuous (straight), glabrous. |
Flowers | sepals (green or yellowish), oblong, 2.2–3 mm, pubescent, (trichomes simple and branched); petals yellow, oblanceolate, 3.5–5 × 1.5–2.5 mm; anthers ovate, 0.4–0.5 mm. |
sepals (erect), oblong or ovate, 2–2.7 mm (lateral pair subsaccate basally), glabrous or abaxially sparsely pilose, (trichomes simple); petals yellow, narrowly obovate, 4–6 × 2–3 mm, (apex emarginate); anthers ovate-subcordate, 0.4–0.5 mm. |
Fruiting pedicels | divaricate-ascending to ascending or suberect, straight, 3–13(–20) mm, pubescent as stem. |
divaricate, straight or curved, (often filiform), 5–18(–23) mm, glabrous. |
Fruits | (often subappressed to rachis), lanceolate to linear-lanceolate or narrowly oblong, slightly twisted or plane, flattened, (6–)9–14(–17) × 2–3.5 mm; valves pubescent, trichomes simple and short-stalked, 2–4-rayed, 0.05–0.3 mm; ovules 28–38(–44) per ovary; style 0.5–1.2(–1.5) mm. |
usually oblong to elliptic, rarely sublinear, plane, flattened, 4–8 × 1.5–2.2 mm; valves glabrous, (obscurely veined); ovules 24–30 per ovary; style 0.5–1 mm. |
Seeds | oblong, 0.9–1.3 × 0.5–0.7 mm. |
(brown), ovoid, 0.9–1.2 × 0.5–0.6 mm. |
2n | = 74. |
= 16. |
Draba aurea |
Draba sibirica |
|
Phenology | Flowering Jun–Aug. | Flowering May–Jul. |
Habitat | Rock outcrops, talus, damp gullies and meadows, subalpine conifer woodlands, alpine slopes and turf, tundra, road banks, river gravel | Wet places on rocky slopes |
Elevation | (0-)700-4200 m ((0-)2300-13800 ft) | 0-300 m (0-1000 ft) |
Distribution |
AK; AZ; CO; ID; MT; NM; NV; SD; UT; WA; WY; AB; BC; MB; NF; NT; NU; ON; QC; SK; YT; Greenland
|
Greenland; Europe (Russia); Asia (Caucasus, Iran, Russian Far East, Siberia, Turkey, central republics) |
Discussion | Draba aurea is extremely variable in plant size, number of cauline leaves, number of bracteate flowers, style length, and fruit size, shape, orientation, twisting, and indumentum. Much of the variation in the number of bracts, style length, fruit twisting, and growth habit occurs in Greenland, where the type specimen was collected and where the species is found near sea level. The highly deviant chromosome counts (e.g., 2n = 40 + 1, 64, 82) listed by R. C. Rollins (1993) and S. I. Warwick and I. A. Al-Shehbaz (2006) are mostly unvouchered and have to be disregarded; counts of 2n = ca. 80 have been re-assigned to Draba glabella. Published (G. A. Mulligan 2002) and unpublished counts made by Mulligan and M. D. Windham from Alaska, British Columbia, Colorado, Quebec, Utah, and Yukon indicate that the most common chromosome number of D. aurea is 2n = 74 (or 72). This suggests that the species is an allopolyploid (hexaploid or higher), incorporating genomes from both euploid and aneuploid lineages (M. A. Beilstein and Windham 2003). Detailed cytological and molecular studies are much needed to fully understand this widely distributed and highly variable species. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Draba sibirica is one of three species of the genus in North America exhibiting malpighiaceous trichomes. It is easily distinguished from others (D. malpighiacea and some plants of D. spectabilis) by a complete absence of cauline leaves. It is found in the flora area in northeastern Greenland. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 294. | FNA vol. 7, p. 337. |
Parent taxa | Brassicaceae > tribe Arabideae > Draba | Brassicaceae > tribe Arabideae > Draba |
Sibling taxa | ||
Synonyms | D. aurea var. aureiformis, D. aurea var. leiocarpa, D. aurea var. luteola, D. aureiformis, D. aureiformis var. leiocarpa, D. bakeri, D. decumbens, D. henneana var. maccallae, D. luteola, D. luteola var. minganensis, D. maccallae, D. minganensis, D. surculifera, D. uber | Lepidium sibiricum, D. gmelinii, D. repens, D. sibirica subsp. arctica |
Name authority | Vahl ex Hornemann: Fors. Oecon. Plantel. ed. 2, 599. (1806) | (Pallas) Thellung: Neue Denkschr. Schweiz. Naturf. Ges. 41: 318. (1907) |
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