The green links below add additional plants to the comparison table. Blue links lead to other Web sites.
enable glossary links

golden Draba, golden Draba whitlow-grass, golden whitlow-grass, twisted-pod Draba

Carolina Draba, Carolina whitlow-grass, Carolina whitlow-mustard, creeping Draba

Habit Perennials; (not cespitose); caudex simple or branched; not scapose. Annuals; scapose or subscapose.
Stems

sometimes branched distally, (0.5–) 1–3.5(–5.2) dm, pubescent throughout, trichomes simple (non-crisped), 0.4–1.3 mm, and 3–6-rayed ones, 0.1–0.5 mm.

(simple or few from, or distal to, base), unbranched, (0.1–)0.3–1.2(–1.6) dm, sparsely to densely pubescent proximally, trichomes 2 (or 3)-rayed, 0.1–0.6 mm, sometimes with simple or spurred ones, to 0.9 mm, distally usually glabrous, rarely with few trichomes.

Basal leaves

rosulate; petiolate;

petiole (distinct or obscure) ciliate, (trichomes simple, to 0.8 mm);

blade oblanceolate to obovate, (0.4–)1–3.7(–5) cm × (1–)2–7(–10) mm, margins entire or denticulate, surfaces pubescent, trichomes stalked, (2–)4–7 (or 8)-rayed, 0.2–0.5(–0.6) mm.

not rosulate;

petiole obscure;

blade elliptic or spatulate to obovate or suborbicular, 0.5–2.3(–3) cm × 1.5–8(–13) mm, margins entire, surfaces pubescent abaxially with stalked, 2–4-rayed trichomes, 0.1–0.5 mm, adaxially with simple trichomes, 0.6–1 mm, with stalked, 2-rayed ones, to 0.7 mm.

Cauline leaves

5–20(–26);

sessile;

blade oblong to lanceolate or ovate, margins entire or dentate, surfaces pubescent as basal, sometimes adaxially with simple trichomes.

usually 0, rarely 1–3;

sessile;

blade similar to basal.

Racemes

(10–)18–52(–72)-flowered, usually bracteate on proximalmost 1–12(–17) flowers, rarely ebracteate, elongated in fruit;

rachis not flexuous, pubescent as stem.

(3–)5–12(–16)-flowered, ebracteate, (subumbellate), not elongated in fruit;

rachis not flexuous, usually glabrous, rarely sparsely pubescent.

Flowers

sepals (green or yellowish), oblong, 2.2–3 mm, pubescent, (trichomes simple and branched);

petals yellow, oblanceolate, 3.5–5 × 1.5–2.5 mm;

anthers ovate, 0.4–0.5 mm.

(late-season ones cleistogamous, apetalous);

sepals oblong, 1.5–2.3 mm, pubescent, (trichomes simple);

petals (rarely absent), white, spatulate, 2–4.5 × 1–1.5 mm;

anthers ovate, 0.4–0.5 mm.

Fruiting pedicels

divaricate-ascending to ascending or suberect, straight, 3–13(–20) mm, pubescent as stem.

horizontal to divaricate-ascending, straight, (1–)2–7(–9) mm, glabrous or glabrate.

Fruits

(often subappressed to rachis), lanceolate to linear-lanceolate or narrowly oblong, slightly twisted or plane, flattened, (6–)9–14(–17) × 2–3.5 mm;

valves pubescent, trichomes simple and short-stalked, 2–4-rayed, 0.05–0.3 mm;

ovules 28–38(–44) per ovary;

style 0.5–1.2(–1.5) mm.

linear to linear-oblong, plane, flattened, (5–)7–16(–20) × 1.2–2.3 mm;

valves glabrous or pubescent, trichomes usually simple and antrorse, 0.1–0.3 mm, rarely with fewer, spurred or 2-rayed ones;

ovules 32–88 per ovary;

style 0.02–0.1 mm.

Seeds

oblong, 0.9–1.3 × 0.5–0.7 mm.

oblong to ovoid, 0.5–0.8 × 0.3–0.5 mm.

2n

= 74.

= 16, 30, 32.

Draba aurea

Draba reptans

Phenology Flowering Jun–Aug. Flowering Feb–Jun(-Aug).
Habitat Rock outcrops, talus, damp gullies and meadows, subalpine conifer woodlands, alpine slopes and turf, tundra, road banks, river gravel Rock outcrops, dry slopes and hillsides, prairies, glades, roadsides, disturbed sites
Elevation (0-)700-4200 m ((0-)2300-13800 ft) 0-3000 m (0-9800 ft)
Distribution
from FNA
AK; AZ; CO; ID; MT; NM; NV; SD; UT; WA; WY; AB; BC; MB; NF; NT; NU; ON; QC; SK; YT; Greenland
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; AZ; CA; CO; GA; IA; ID; IL; KS; MA; MI; MN; MO; MT; NC; ND; NE; NM; NY; OH; OK; OR; PA; SC; SD; TN; TX; UT; WA; WI; WY; AB; BC; MB; ON; SK
[WildflowerSearch map]
[BONAP county map]
Discussion

Draba aurea is extremely variable in plant size, number of cauline leaves, number of bracteate flowers, style length, and fruit size, shape, orientation, twisting, and indumentum. Much of the variation in the number of bracts, style length, fruit twisting, and growth habit occurs in Greenland, where the type specimen was collected and where the species is found near sea level.

The highly deviant chromosome counts (e.g., 2n = 40 + 1, 64, 82) listed by R. C. Rollins (1993) and S. I. Warwick and I. A. Al-Shehbaz (2006) are mostly unvouchered and have to be disregarded; counts of 2n = ca. 80 have been re-assigned to Draba glabella. Published (G. A. Mulligan 2002) and unpublished counts made by Mulligan and M. D. Windham from Alaska, British Columbia, Colorado, Quebec, Utah, and Yukon indicate that the most common chromosome number of D. aurea is 2n = 74 (or 72). This suggests that the species is an allopolyploid (hexaploid or higher), incorporating genomes from both euploid and aneuploid lineages (M. A. Beilstein and Windham 2003). Detailed cytological and molecular studies are much needed to fully understand this widely distributed and highly variable species.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Draba reptans is often confused with D. cuneifolia, but the two are easily separated. The rachises and pedicels of D. reptans are usually glabrous (rarely with a few isolated trichomes); those of D. cuneifolia are always densely pubescent. Interestingly, both species show parallel variations in chromosome number; it is currently unclear whether this variation is real or the result of misidentified specimens and/or erroneous counts.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 294. FNA vol. 7, p. 333.
Parent taxa Brassicaceae > tribe Arabideae > Draba Brassicaceae > tribe Arabideae > Draba
Sibling taxa
D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. cyclomorpha, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. helleriana, D. hitchcockii, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis
D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. cyclomorpha, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. helleriana, D. hitchcockii, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis
Synonyms D. aurea var. aureiformis, D. aurea var. leiocarpa, D. aurea var. luteola, D. aureiformis, D. aureiformis var. leiocarpa, D. bakeri, D. decumbens, D. henneana var. maccallae, D. luteola, D. luteola var. minganensis, D. maccallae, D. minganensis, D. surculifera, D. uber Arabis reptans, D. caroliniana, D. caroliniana var. dolichocarpa, D. caroliniana var. hunteri, D. caroliniana var. micrantha, D. caroliniana subsp. stellifera, D. caroliniana var. umbellata, D. coloradensis, D. hispidula, D. micrantha, D. reptans var. micrantha, D. reptans subsp. stellifera, D. reptans var. stellifera, D. umbellata, Tomostima caroliniana, Tomostima hispidula, Tomostima micrantha
Name authority Vahl ex Hornemann: Fors. Oecon. Plantel. ed. 2, 599. (1806) (Lamarck) Fernald: Rhodora 36: 368. (1934)
Web links