Flora of North America species comparison

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Draba aurea

Draba arabisans

golden Draba, golden Draba whitlow-grass, golden whitlow-grass, twisted-pod Draba

rock Draba, rock whitlow-mustard
Habit Perennials; (not cespitose); caudex simple or branched; not scapose. Perennials; (sometimes cespitose); caudex simple or branched; not scapose.
Stems

sometimes branched distally, (0.5–) 1–3.5(–5.2) dm, pubescent throughout, trichomes simple (non-crisped), 0.4–1.3 mm, and 3–6-rayed ones, 0.1–0.5 mm.

branched or unbranched, (0.5–)0.7–3(–4.5) dm, pubescent proximally, glabrous or pubescent distally, trichomes sessile (non-crisped), 4- or 5-rayed, stellate, or subpectinate, 0.07–0.4 mm.
Basal leaves

rosulate; petiolate;

petiole (distinct or obscure) ciliate, (trichomes simple, to 0.8 mm);

blade oblanceolate to obovate, (0.4–)1–3.7(–5) cm × (1–)2–7(–10) mm, margins entire or denticulate, surfaces pubescent, trichomes stalked, (2–)4–7 (or 8)-rayed, 0.2–0.5(–0.6) mm.

rosulate; petiolate;

petiole base and margin sparsely ciliate, (trichomes simple and 2-rayed, 0.2–0.8 mm);

blade oblanceolate to spatulate or linear-oblanceolate, 1–3.8(–5) cm × 2–5(–8) mm, margins dentate or denticulate, surfaces pubescent with sessile, non-crisped, 4- or 5-rayed, stellate trichomes, 0.07–0.5mm, (some rays branched, appearing 2–12-rayed, midvein obscure abaxially).
Cauline leaves

5–20(–26);

sessile;

blade oblong to lanceolate or ovate, margins entire or dentate, surfaces pubescent as basal, sometimes adaxially with simple trichomes.

(2 or) 3–10(–13);

sessile;

blade ovate to oblong, margins dentate to subentire, surfaces pubescent as basal.
Racemes

(10–)18–52(–72)-flowered, usually bracteate on proximalmost 1–12(–17) flowers, rarely ebracteate, elongated in fruit;

rachis not flexuous, pubescent as stem.

5–18(–25)-flowered, ebracteate, elongated in fruit;

rachis not flexuous, glabrous or pubescent as stem.
Flowers

sepals (green or yellowish), oblong, 2.2–3 mm, pubescent, (trichomes simple and branched);

petals yellow, oblanceolate, 3.5–5 × 1.5–2.5 mm;

anthers ovate, 0.4–0.5 mm.

sepals oblong, 1.7–2.5 mm, pubescent, trichomes short-stalked, 3- or 4-rayed;

petals white, broadly obovate, 3.5–5 × 1.5–2.5 mm;

anthers ovate, 0.4–0.5 mm.
Fruiting pedicels

divaricate-ascending to ascending or suberect, straight, 3–13(–20) mm, pubescent as stem.

divaricate-ascending to suberect, straight or slightly curved, (3–)4–10(–13) mm, glabrous or pubescent as stem.
Fruits

(often subappressed to rachis), lanceolate to linear-lanceolate or narrowly oblong, slightly twisted or plane, flattened, (6–)9–14(–17) × 2–3.5 mm;

valves pubescent, trichomes simple and short-stalked, 2–4-rayed, 0.05–0.3 mm;

ovules 28–38(–44) per ovary;

style 0.5–1.2(–1.5) mm.

ovate to lanceolate or linear-lanceolate, usually slightly twisted, rarely plane, flattened, 5–11(–14) × 2–3.5 mm;

valves usually glabrous, rarely pubescent, trichomes 3- or 4-rayed, 0.07–0.2 mm;

ovules 20–28(–32) per ovary;

style (0.2–)0.4–1 mm.
Seeds

oblong, 0.9–1.3 × 0.5–0.7 mm.

oblong, 0.8–1.1 × 0.6–0.8 mm.
2n

= 74.

= 96.

Draba aurea

Draba arabisans
Phenology Flowering Jun–Aug. Flowering Jun–Aug.
Habitat Rock outcrops, talus, damp gullies and meadows, subalpine conifer woodlands, alpine slopes and turf, tundra, road banks, river gravel Rock outcrops and talus (usually limestone or dolomite), open woods, rocky shores
Elevation (0-)700-4200 m ((0-)2300-13800 ft) 0-300 m (0-1000 ft)
Distribution
from FNA
AK; AZ; CO; ID; MT; NM; NV; SD; UT; WA; WY; AB; BC; MB; NF; NT; NU; ON; QC; SK; YT; Greenland
[WildflowerSearch map]
[BONAP county map]
from FNA
ME; MI; MN; NY; VT; WI; MB; NB; NF; NS; ON; QC
[WildflowerSearch map]
[BONAP county map]
Discussion

Draba aurea is extremely variable in plant size, number of cauline leaves, number of bracteate flowers, style length, and fruit size, shape, orientation, twisting, and indumentum. Much of the variation in the number of bracts, style length, fruit twisting, and growth habit occurs in Greenland, where the type specimen was collected and where the species is found near sea level.

The highly deviant chromosome counts (e.g., 2n = 40 + 1, 64, 82) listed by R. C. Rollins (1993) and S. I. Warwick and I. A. Al-Shehbaz (2006) are mostly unvouchered and have to be disregarded; counts of 2n = ca. 80 have been re-assigned to Draba glabella. Published (G. A. Mulligan 2002) and unpublished counts made by Mulligan and M. D. Windham from Alaska, British Columbia, Colorado, Quebec, Utah, and Yukon indicate that the most common chromosome number of D. aurea is 2n = 74 (or 72). This suggests that the species is an allopolyploid (hexaploid or higher), incorporating genomes from both euploid and aneuploid lineages (M. A. Beilstein and Windham 2003). Detailed cytological and molecular studies are much needed to fully understand this widely distributed and highly variable species.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Draba arabisans is often confused with D. glabella (2n = 80); it is easily distinguished by having sessile (versus short-stalked) stellate trichomes on abaxial leaf blade surfaces and longer styles (0.2–)0.4–1 [versus 0.1–0.2(–0.4)] mm. Both species have twisted and plane fruits; D. arabisans has mostly twisted fruits; D. glabella has predominantly plane fruits. The complete absence of simple trichomes proximally on stems of D. arabisans can be a useful feature separating it from D. glabella, which often has at least some simple trichomes on the proximal half of the stem.

Draba arabis Persoon is an illegitimate name, sometimes found in synonymy with D. arabisans.

(Discussion copyrighted by Flora of North America; reprinted with permission.)
Source FNA vol. 7, p. 294. FNA vol. 7, p. 291.
Parent taxa Brassicaceae > tribe Arabideae > Draba Brassicaceae > tribe Arabideae > Draba
Sibling taxa
D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. cyclomorpha, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. helleriana, D. hitchcockii, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis
D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. cyclomorpha, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. helleriana, D. hitchcockii, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis
Synonyms D. aurea var. aureiformis, D. aurea var. leiocarpa, D. aurea var. luteola, D. aureiformis, D. aureiformis var. leiocarpa, D. bakeri, D. decumbens, D. henneana var. maccallae, D. luteola, D. luteola var. minganensis, D. maccallae, D. minganensis, D. surculifera, D. uber D. arabisans var. superiorensis, D. incana var. arabisans, D. incana var. glabriuscula
Name authority Vahl ex Hornemann: Fors. Oecon. Plantel. ed. 2, 599. (1806) Michaux: Fl. Bor.-Amer. 2: 28. (1803)
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Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).



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