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fellfield whitlowgrass

Smith's Draba

Habit Perennials; (cespitose); caudex branched (sometimes with persistent leaf remains); sometimes scapose. Perennials; (matted); caudex branched (with persistent leaf remains, branches slender, prostrate, sometimes terminating in sterile rosettes); not scapose.
Stems

unbranched, 0.3–1.4 dm, pubescent throughout, trichomes simple, 0.2–1 mm, and 2–6-rayed, 0.05–0.3 mm.

usually unbranched, (0.5–)1–2.4(–3) dm, pubescent throughout, trichomes short-stalked, subdendritic, 4–12-rayed, 0.05–0.2 mm.

Basal leaves

rosulate;

petiolate;

petiole base and margin ciliate, (trichomes mostly simple, 0.2–1 mm);

blade oblanceolate to narrowly obovate, 0.4–1.7 cm × 1.5–6 mm, margins entire or with 1–3 teeth on each side, surfaces pubescent with primarily simple trichomes, 0.3–1mm, abaxially mixed with stalked, 4–10-rayed ones, 0.1–0.2 mm.

rosulate;

petiole (obscure), usually not ciliate, rarely pubescent proximally, (trichomes simple or spurred, to 0.25 mm);

blade obovate to narrowly oblanceolate, 0.5–1.3(–2.5) cm × (1–)2–4(–7) mm, margins usually entire, rarely denticulate, surfaces pubescent with short-stalked, subdendritic, 5–12-rayed trichomes, 0.05–0.3 mm.

Cauline leaves

0 or 1;

sessile;

blade ovate to oblong, margins entire or toothed, surfaces pubescent as basal.

(2 or) 3–8;

sessile;

blade oblong to lanceolate, margins usually entire, rarely denticulate, surfaces pubescent as basal, or, rarely, adaxially with long-stalked, 2–5-rayed trichomes.

Racemes

2–14(–17)-flowered, ebracteate, elongated in fruit;

rachis not flexuous, pubescent as stem.

12–28-flowered, ebracteate, elongated in fruit;

rachis not flexuous, pubescent as stem.

Flowers

sepals ovate, 1.8–2.5 mm, pubescent, (trichomes simple and short-stalked, 2–4-rayed);

petals white, spatulate, 3–4 × 1.5–2 mm;

anthers ovate, 0.3–0.4 mm.

sepals ovate, 2–2.5 mm, pubescent;

petals white, spatulate, 4–6 × 2–3 mm;

anthers ovate, 0.5–0.6 mm.

Fruiting pedicels

suberect to ascending, straight, 1–5(–7) mm, pubescent, trichomes simple and 2–4-rayed.

divaricate, straight, (3–)5–10(–13) mm, pubescent as stem.

Fruits

oblong, plane, flattened, 4–9 × 2–3.5 mm;

valves pubescent, trichomes short-stalked, 2–4-rayed, 0.1–0.5 mm, sometimes with simple ones;

ovules 20–32 per ovary;

style 0.1–0.4 mm (stigma distinctly wider than style).

ovate-lanceolate, twisted, flattened, 5–9 × 2–3 mm;

valves pubescent, trichomes simple and short-stalked, 2–4-rayed, 0.05–0.2 mm;

ovules 16–20 per ovary;

style (0.7–)1–2(–2.3) mm.

Seeds

ovoid, 0.7–1 × 0.5–0.6 mm.

oblong, 0.8–1.2 × 0.5–0.7 mm.

2n

= 48.

= 32.

Draba arctogena

Draba smithii

Phenology Flowering Jun–Jul. Flowering May–Jul.
Habitat Scree, sandy hills, gravel flats Igneous outcrops and rocky slopes in mixed conifer, mountain shrub, and (rarely) sagebrush communities
Elevation 0-450 m (0-1500 ft) 2300-3700 m (7500-12100 ft)
Distribution
from FNA
NT; NU; Greenland
[BONAP county map]
from FNA
CO
[BONAP county map]
Discussion

Draba arctogena is sometimes confused with D. norvegica, from which it differs by having less elongated fruiting racemes, abaxial leaf surfaces with 4–10-rayed (versus cruciform or, rarely, 2–6-rayed) trichomes, and fruits with 2–4-rayed trichomes, 0.1–0.5 mm, (versus glabrous or with simple and 2–4-rayed trichomes, 0.05–0.25 mm). The limits of D. arctica, D. arctogena, D. cinerea, and D. oblongata need to be critically evaluated in light of molecular and cytological data.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Draba smithii is a distinctive species known only from Archuleta, Las Animas, and Mineral counties in southern Colorado.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 292. FNA vol. 7, p. 338.
Parent taxa Brassicaceae > tribe Arabideae > Draba Brassicaceae > tribe Arabideae > Draba
Sibling taxa
D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. cyclomorpha, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. helleriana, D. hitchcockii, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis
D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. cyclomorpha, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. helleriana, D. hitchcockii, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis
Synonyms D. groenlandica var. arctogena, D. cinerea var. arctogena
Name authority (E. Ekman) E. Ekman: Svensk Bot. Tidskr. 25: 492. (1932) Gilg & O. E. Schulz: in H. G. A. Engler, Pflanzenr. 89[IV,105]: 177. (1927)
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