Draba arctogena |
Draba porsildii |
|
---|---|---|
fellfield whitlowgrass |
Porsild's Draba, Porsild's whitlow-grass |
|
Habit | Perennials; (cespitose); caudex branched (sometimes with persistent leaf remains); sometimes scapose. | Perennials; (cespitose); caudex branched (with persistent leaf remains); often scapose. |
Stems | unbranched, 0.3–1.4 dm, pubescent throughout, trichomes simple, 0.2–1 mm, and 2–6-rayed, 0.05–0.3 mm. |
unbranched, 0.2–1.2 dm, sparsely pubescent proximally, trichomes short-stalked, 3–5- rayed, (non-crisped), 0.2–0.4 mm, glabrous distally. |
Basal leaves | rosulate; petiolate; petiole base and margin ciliate, (trichomes mostly simple, 0.2–1 mm); blade oblanceolate to narrowly obovate, 0.4–1.7 cm × 1.5–6 mm, margins entire or with 1–3 teeth on each side, surfaces pubescent with primarily simple trichomes, 0.3–1mm, abaxially mixed with stalked, 4–10-rayed ones, 0.1–0.2 mm. |
rosulate; subsessile or shortly petiolate; petiole base and margin ciliate proximally, (trichomes simple, subsetiform, stiff, 0.3–0.9 mm); blade oblanceolate to obovate, 0.4–1.5 cm × 1–3 mm, margins entire, surfaces pubescent, abaxially with stalked, 6–12-rayed, (non-crisped) trichomes, 0.2–0.5 mm, (midvein obscure), adaxially similar, or with simple trichomes on proximal 1/2. |
Cauline leaves | 0 or 1; sessile; blade ovate to oblong, margins entire or toothed, surfaces pubescent as basal. |
0 or 1; sessile; blade oblong to lanceolate, margins entire, surfaces pubescent as basal. |
Racemes | 2–14(–17)-flowered, ebracteate, elongated in fruit; rachis not flexuous, pubescent as stem. |
3–13-flowered, ebracteate, elongated in fruit; rachis not flexuous, glabrous. |
Flowers | sepals ovate, 1.8–2.5 mm, pubescent, (trichomes simple and short-stalked, 2–4-rayed); petals white, spatulate, 3–4 × 1.5–2 mm; anthers ovate, 0.3–0.4 mm. |
sepals ovate, 1.5–2 mm, pubescent, (trichomes simple and 2-rayed); petals white, spatulate, 2–4 × 1–1.7 mm; anthers ovate, ca. 0.2 mm. |
Fruiting pedicels | suberect to ascending, straight, 1–5(–7) mm, pubescent, trichomes simple and 2–4-rayed. |
divaricate-ascending, straight, 1–4(–8) mm, glabrous. |
Fruits | oblong, plane, flattened, 4–9 × 2–3.5 mm; valves pubescent, trichomes short-stalked, 2–4-rayed, 0.1–0.5 mm, sometimes with simple ones; ovules 20–32 per ovary; style 0.1–0.4 mm (stigma distinctly wider than style). |
ovate to oblong, plane, flattened, 4–7.5 × 1.7–3 mm; valves glabrous; ovules 12–16 per ovary; style 0.1–0.3(–0.5) mm. |
Seeds | ovoid, 0.7–1 × 0.5–0.6 mm. |
ovoid, 0.7–1 × 0.5–0.6 mm. |
2n | = 48. |
= 32. |
Draba arctogena |
Draba porsildii |
|
Phenology | Flowering Jun–Jul. | Flowering Jun–Jul. |
Habitat | Scree, sandy hills, gravel flats | Rock outcrops, talus, meadows, gravel slopes |
Elevation | 0-450 m (0-1500 ft) | 600-3000 m (2000-9800 ft) |
Distribution |
NT; NU; Greenland |
AK; MT; WY; AB; YT |
Discussion | Draba arctogena is sometimes confused with D. norvegica, from which it differs by having less elongated fruiting racemes, abaxial leaf surfaces with 4–10-rayed (versus cruciform or, rarely, 2–6-rayed) trichomes, and fruits with 2–4-rayed trichomes, 0.1–0.5 mm, (versus glabrous or with simple and 2–4-rayed trichomes, 0.05–0.25 mm). The limits of D. arctica, D. arctogena, D. cinerea, and D. oblongata need to be critically evaluated in light of molecular and cytological data. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
According to H. H. Grundt et al. (2004), tetraploid Draba porsildii probably originated from a diploid ancestor such as D. lonchocarpa. The Wyoming plants are a close match to the type of the species, though they have distinctly shorter pedicels. Some of the Alaskan plants are more problematic; they have finer and more branched trichomes, larger flowers, longer pedicels, and longer styles than the type. It is likely that they belong to another species; detailed molecular and cytological studies on this complex are needed before any meaningful conclusions are reached. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 292. | FNA vol. 7, p. 331. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | D. groenlandica var. arctogena, D. cinerea var. arctogena | D. nivalis var. brevicula, D. porsildii var. brevicula |
Name authority | (E. Ekman) E. Ekman: Svensk Bot. Tidskr. 25: 492. (1932) | G. A. Mulligan: Canad. J. Bot. 52: 1795, fig. 8. (1974) |
Web links |