Draba alpina |
Draba ventosa |
|
---|---|---|
alpine Draba |
wind river Draba, wind river whitlow-grass |
|
Habit | Perennials; (cespitose); caudex branched (covered with persistent leaves or leaf remains); scapose. | Perennials; (cespitose); caudex branched (covered with persistent leaves, branches creeping, sometimes terminating in sterile rosettes); scapose. |
Stems | unbranched, (0.3–)0.5–1.7(–2.8) dm, pubescent throughout, trichomes simple and 2-rayed, 0.3–0.8 mm, with 3–5-rayed ones, 0.1–0.3 mm. |
unbranched, (0.1–)0.2–0.4(–0.6) dm, densely pubescent throughout, trichomes 2–6-rayed, 0.1–0.6 mm. |
Basal leaves | rosulate; petiolate; petiole base (not thickened), ciliate, (trichomes simple, 0.3–1 mm); blade oblanceolate to obovate or lanceolate to oblong, 0.8–3(–4.5) cm × 2.5–6(–9) mm, margins entire, surfaces abaxially pubescent with stalked, 2–4-rayed trichomes, 0.1–0.5 mm, with simple ones (midvein obscure, not thickened), adaxially glabrous or sparsely pubescent with simple and stalked, 2–4-rayed trichomes. |
(imbricate); rosulate; subsessile; petiole base and margin not ciliate; blade obovate to oblanceolate, (0.4–)0.5–1 cm × 1.5–4.5 mm, margins entire, surfaces densely pubescent with stalked, 2–6-rayed trichomes, (0.1–)0.2–0.9 mm. |
Cauline leaves | 0. |
0. |
Racemes | 6–18-flowered, ebracteate, considerably elongated in fruit; rachis not flexuous, pubescent as stem. |
5–10(–16)-flowered, ebracteate, slightly elongated in fruit; rachis not flexuous, densely pubescent as stem. |
Flowers | sepals (purplish tinged), narrowly ovate, 2.5–3 mm, pubescent, (trichomes simple and fewer, stalked, 2-rayed); petals bright yellow, narrowly obovate, 3.5–5 × 1.7–2.5 mm; anthers ovate, 0.3–0.4 mm. |
sepals broadly ovate, 2–2.5 mm, pubescent, (trichomes short-stalked, 2–6-rayed); petals yellow, obovate, 3.5–5.5 × 1.5–3 mm; anthers oblong, 0.5–0.6 mm. |
Fruiting pedicels | ascending to divaricate-ascending, straight or, sometimes, slightly curved upwards, 4–14(–30) mm, pubescent, trichomes simple and 2–4-rayed. |
horizontal to divaricate-ascending, straight or slightly curved upward, 2–7(–9) mm, densely pubescent, trichomes 2–6-rayed, (0.1–0.6 mm). |
Fruits | elliptic, plane, flattened, 6–10 × 2–3 mm; valves glabrous or glabrescent, trichomes simple, (not confined to replum); ovules 12–24 per ovary; style 0.2–0.3 mm (stigma about as wide as style). |
suborbicular to broadly ovate, plane, inflated basally, flattened distally, 4–7.5(–9) × 3.5–5 mm; valves densely pubescent, trichomes short-stalked, 2–6-rayed, 0.15–0.5 mm; ovules 8–12 per ovary; style (0.5–)0.7–1.4 mm. |
Seeds | (pale brown), ovoid, 0.9–1.3 × 0.6–0.9 mm. |
oblong, 1.4–1.9 × 0.9–1.2 mm. |
2n | = 80. |
= 36. |
Draba alpina |
Draba ventosa |
|
Phenology | Flowering Jun–Aug. | Flowering Jul–Aug. |
Habitat | Moist tundra and ridges, sand and gravel flats or beaches | Talus slopes and alpine tundra |
Elevation | 0-1000 m (0-3300 ft) | 2000-4000 m (6600-13100 ft) |
Distribution |
LB; MB; NU; ON; QC; Greenland; Europe (Finland, Norway, Russia, Sweden)
|
CO; UT; WY; AB; BC; YT |
Discussion | The synonymy above includes two North American names overlooked by C. L. Hitchcock (1941) and R. C. Rollins (1993). Draba alpina was broadly delimited by O. E. Schulz (1927) and included 17 varieties, some of which (e.g., corymbosa, oxycarpa, pilosa) are recognized herein as distinct species. The name D. alpina was so misapplied that it was used for any circumpolar or alpine, scapose, yellow-flowered, perennial Draba. Various chromosome numbers (e.g., 2n = 64, 80, 112, 120; S. I. Warwick and I. A. Al-Shehbaz 2006) have been reported for the species. As circumscribed here, it has the narrow distribution outlined above and includes plants with 2n = 80. Reports of the species from Alaska, Canadian Northwest Territories and Yukon, Siberia, eastern Asia, Russian Far East, and the Central Asian republics are either suspect or very unlikely. The entire D. alpina complex (including the above three species, D. glacialis Adams, D. macounii, etc.) is in need of critical molecular, cytological, and morphological study. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
G. A. Mulligan (1971b) first reported apomixis in Draba ventosa based on studies of Canadian populations. This has now been confirmed in one of the southernmost populations (Duchesne County, Utah; M. D. Windham, unpubl.) as well. The species is easily overlooked and the large geographic gap between the Canadian and United States populations is likely to be narrowed or eliminated by additional collecting in western Montana. The limits of this species were expanded by C. L. Hitchcock (1941) to include D. ruaxes, but there are clear differences between them that support their recognition as distinct species. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 290. | FNA vol. 7, p. 344. |
Parent taxa | Brassicaceae > tribe Arabideae > Draba | Brassicaceae > tribe Arabideae > Draba |
Sibling taxa | ||
Synonyms | D. alpina var. hydeana, D. alpina var. inflatisiliqua | |
Name authority | Linnaeus: Sp. Pl. 2: 642. (1753) | A. Gray: Amer. Naturalist 8: 212. (1874) |
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