Draba alpina |
Draba standleyi |
|
---|---|---|
alpine Draba |
Standley's Draba |
|
Habit | Perennials; (cespitose); caudex branched (covered with persistent leaves or leaf remains); scapose. | Perennials; (densely pulvinate); caudex branched (with persistent, thickened petioles, branches compact); not scapose. |
Stems | unbranched, (0.3–)0.5–1.7(–2.8) dm, pubescent throughout, trichomes simple and 2-rayed, 0.3–0.8 mm, with 3–5-rayed ones, 0.1–0.3 mm. |
unbranched, 0.3–1.3(–1.7) dm, usually glabrous throughout or sparsely pubescent proximally, rarely sparsely pubescent distally, trichomes simple and 2-rayed, 0.1–0.7 mm. |
Basal leaves | rosulate; petiolate; petiole base (not thickened), ciliate, (trichomes simple, 0.3–1 mm); blade oblanceolate to obovate or lanceolate to oblong, 0.8–3(–4.5) cm × 2.5–6(–9) mm, margins entire, surfaces abaxially pubescent with stalked, 2–4-rayed trichomes, 0.1–0.5 mm, with simple ones (midvein obscure, not thickened), adaxially glabrous or sparsely pubescent with simple and stalked, 2–4-rayed trichomes. |
(not imbricate); rosulate; petiolate; petiole ciliate, (trichomes usually simple, rarely 2-rayed, not setiform); blade narrowly oblanceolate to linear-lanceolate, (strongly differentiated into blade and petiole), (1.2–)1.8–6(–8.5) cm × 1–5(–7) mm, margins entire or sparsely denticulate, (ciliate as petiole; midvein not prominent), surfaces glabrous or pubescent, usually with simple trichomes 0.1–0.8 mm, rarely 2-rayed. |
Cauline leaves | 0. |
1–8; sessile; blade lanceolate to narrowly oblong, margins usually entire, surfaces pubescent as basal. |
Racemes | 6–18-flowered, ebracteate, considerably elongated in fruit; rachis not flexuous, pubescent as stem. |
5–17(–23)-flowered, ebracteate, elongated in fruit; rachis not flexuous, usually glabrous, rarely pubescent, trichomes simple and 2-rayed, (non-crisped). |
Flowers | sepals (purplish tinged), narrowly ovate, 2.5–3 mm, pubescent, (trichomes simple and fewer, stalked, 2-rayed); petals bright yellow, narrowly obovate, 3.5–5 × 1.7–2.5 mm; anthers ovate, 0.3–0.4 mm. |
sepals ovate, 2–2.5 mm, pubescent, (trichomes simple); petals yellow, oblanceolate, 4–6 × 1.5–2 mm; anthers oblong, 0.6–0.8 mm. |
Fruiting pedicels | ascending to divaricate-ascending, straight or, sometimes, slightly curved upwards, 4–14(–30) mm, pubescent, trichomes simple and 2–4-rayed. |
divaricate-ascending or ascending, straight, 3–9(–13) mm, usually glabrous, rarely sparsely pubescent, trichomes simple. |
Fruits | elliptic, plane, flattened, 6–10 × 2–3 mm; valves glabrous or glabrescent, trichomes simple, (not confined to replum); ovules 12–24 per ovary; style 0.2–0.3 mm (stigma about as wide as style). |
linear-elliptic to elliptic, twisted or plane, flattened, 5–10(–13) × 1.5–2.5 mm; valves usually glabrous, rarely puberulent, trichomes simple, 0.05–0.1 mm; ovules 12–24 per ovary; style 0.7–1.4(–1.8) mm. |
Seeds | (pale brown), ovoid, 0.9–1.3 × 0.6–0.9 mm. |
ovoid, 1–1.2 × 0.7–0.8 mm. |
2n | = 80. |
|
Draba alpina |
Draba standleyi |
|
Phenology | Flowering Jun–Aug. | Flowering Jun–Aug. |
Habitat | Moist tundra and ridges, sand and gravel flats or beaches | Igneous rock outcrops, stabilized talus slopes |
Elevation | 0-1000 m (0-3300 ft) | 1800-3100 m (5900-10200 ft) |
Distribution |
LB; MB; NU; ON; QC; Greenland; Europe (Finland, Norway, Russia, Sweden)
|
AZ; NM; TX |
Discussion | The synonymy above includes two North American names overlooked by C. L. Hitchcock (1941) and R. C. Rollins (1993). Draba alpina was broadly delimited by O. E. Schulz (1927) and included 17 varieties, some of which (e.g., corymbosa, oxycarpa, pilosa) are recognized herein as distinct species. The name D. alpina was so misapplied that it was used for any circumpolar or alpine, scapose, yellow-flowered, perennial Draba. Various chromosome numbers (e.g., 2n = 64, 80, 112, 120; S. I. Warwick and I. A. Al-Shehbaz 2006) have been reported for the species. As circumscribed here, it has the narrow distribution outlined above and includes plants with 2n = 80. Reports of the species from Alaska, Canadian Northwest Territories and Yukon, Siberia, eastern Asia, Russian Far East, and the Central Asian republics are either suspect or very unlikely. The entire D. alpina complex (including the above three species, D. glacialis Adams, D. macounii, etc.) is in need of critical molecular, cytological, and morphological study. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Draba standleyi is a distinctive species that is sporadically distributed in the mountains of southwestern United States. It is known from the Chiricahua Mountains (Cochise County, southeastern Arizona), the Organ Mountains and Black Range (Dona Ana and Sierra counties, south-central New Mexico), and the Davis Mountains (Jeff Davis County, western Texas). It has not been reported from Mexico, though it is very likely to occur there. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 290. | FNA vol. 7, p. 340. |
Parent taxa | Brassicaceae > tribe Arabideae > Draba | Brassicaceae > tribe Arabideae > Draba |
Sibling taxa | ||
Synonyms | D. alpina var. hydeana, D. alpina var. inflatisiliqua | D. gilgiana, D. chrysantha var. gilgiana |
Name authority | Linnaeus: Sp. Pl. 2: 642. (1753) | J. F. Macbride & Payson: Ann. Missouri Bot. Gard. 5: 150. (1918) |
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