Draba alpina |
Draba saxosa |
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alpine Draba |
Southern California Rock Draba |
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Habit | Perennials; (cespitose); caudex branched (covered with persistent leaves or leaf remains); scapose. | Perennials; (cespitose); caudex branched (covered with persistent leaves); scapose. |
Stems | unbranched, (0.3–)0.5–1.7(–2.8) dm, pubescent throughout, trichomes simple and 2-rayed, 0.3–0.8 mm, with 3–5-rayed ones, 0.1–0.3 mm. |
unbranched, 0.5–1.5 dm, pubescent throughout, trichomes simple and long-stalked, 2-rayed, 0.3–1 mm, with smaller, 2–4-rayed ones, 0.1–0.5 mm. |
Basal leaves | rosulate; petiolate; petiole base (not thickened), ciliate, (trichomes simple, 0.3–1 mm); blade oblanceolate to obovate or lanceolate to oblong, 0.8–3(–4.5) cm × 2.5–6(–9) mm, margins entire, surfaces abaxially pubescent with stalked, 2–4-rayed trichomes, 0.1–0.5 mm, with simple ones (midvein obscure, not thickened), adaxially glabrous or sparsely pubescent with simple and stalked, 2–4-rayed trichomes. |
rosulate; petiolate; petiole base and margin ciliate, (trichomes simple, 0.5–1.2 mm); blade oblanceolate, 1–3.1 cm × 2–7 mm, margins entire, surfaces densely pubescent, abaxially with stalked, (2–)4-rayed trichomes, 0.2–0.8 mm, adaxially mostly with long-stalked, 2-rayed trichomes, 0.6–1.3 mm, sometimes with simple and 3- or 4-rayed ones. |
Cauline leaves | 0. |
0 (or 1); sessile; blade oblong, margins entire, surfaces pubescent as basal. |
Racemes | 6–18-flowered, ebracteate, considerably elongated in fruit; rachis not flexuous, pubescent as stem. |
12–43-flowered, ebracteate, elongated in fruit; rachis not flexuous, pubescent as stem. |
Flowers | sepals (purplish tinged), narrowly ovate, 2.5–3 mm, pubescent, (trichomes simple and fewer, stalked, 2-rayed); petals bright yellow, narrowly obovate, 3.5–5 × 1.7–2.5 mm; anthers ovate, 0.3–0.4 mm. |
sepals oblong, 2–2.5 mm, pubescent, (trichomes short-stalked, 2–4-rayed); petals yellow, linear-oblanceolate, 3–3.5 × 0.5–1 mm; anthers oblong, 0.5–0.6 mm, (exserted). |
Fruiting pedicels | ascending to divaricate-ascending, straight or, sometimes, slightly curved upwards, 4–14(–30) mm, pubescent, trichomes simple and 2–4-rayed. |
divaricate to ascending, straight or curved, (6–)10–17 mm, pubescent as stem. |
Fruits | elliptic, plane, flattened, 6–10 × 2–3 mm; valves glabrous or glabrescent, trichomes simple, (not confined to replum); ovules 12–24 per ovary; style 0.2–0.3 mm (stigma about as wide as style). |
oblong to linear-oblong, slightly twisted or plane, flattened, 6–10 × 2.5–4 mm; valves pubescent, trichomes short-stalked, cruciform, 0.07–0.25 mm, sometimes with fewer, 2- or 3-rayed ones; ovules 8–24 per ovary; style (0.8–)1.2–3.5 mm. |
Seeds | (pale brown), ovoid, 0.9–1.3 × 0.6–0.9 mm. |
oblong, 1–1.2 × 0.6–0.8 mm. |
2n | = 80. |
|
Draba alpina |
Draba saxosa |
|
Phenology | Flowering Jun–Aug. | Flowering Jun–Jul. |
Habitat | Moist tundra and ridges, sand and gravel flats or beaches | Among rocks |
Elevation | 0-1000 m (0-3300 ft) | 2600-3300 m (8500-10800 ft) |
Distribution |
LB; MB; NU; ON; QC; Greenland; Europe (Finland, Norway, Russia, Sweden)
|
CA |
Discussion | The synonymy above includes two North American names overlooked by C. L. Hitchcock (1941) and R. C. Rollins (1993). Draba alpina was broadly delimited by O. E. Schulz (1927) and included 17 varieties, some of which (e.g., corymbosa, oxycarpa, pilosa) are recognized herein as distinct species. The name D. alpina was so misapplied that it was used for any circumpolar or alpine, scapose, yellow-flowered, perennial Draba. Various chromosome numbers (e.g., 2n = 64, 80, 112, 120; S. I. Warwick and I. A. Al-Shehbaz 2006) have been reported for the species. As circumscribed here, it has the narrow distribution outlined above and includes plants with 2n = 80. Reports of the species from Alaska, Canadian Northwest Territories and Yukon, Siberia, eastern Asia, Russian Far East, and the Central Asian republics are either suspect or very unlikely. The entire D. alpina complex (including the above three species, D. glacialis Adams, D. macounii, etc.) is in need of critical molecular, cytological, and morphological study. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Draba saxosa is related to D. corrugata and is often considered a variety of that species. The two taxa are easily distinguished (I. A. Al-Shehbaz and M. D. Windham 2007) and the available data support their recognition as distinct species. Draba saxosa is restricted to the San Jacinto and Santa Rosa mountains in southern California (Riverside and San Diego counties). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 290. | FNA vol. 7, p. 335. |
Parent taxa | Brassicaceae > tribe Arabideae > Draba | Brassicaceae > tribe Arabideae > Draba |
Sibling taxa | ||
Synonyms | D. alpina var. hydeana, D. alpina var. inflatisiliqua | D. corrugata var. saxosa |
Name authority | Linnaeus: Sp. Pl. 2: 642. (1753) | Davidson: Bull. S. Calif. Acad. Sci. 19: 11. (1920) |
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