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alpine Draba

Habit Perennials; (cespitose); caudex branched (covered with persistent leaves or leaf remains); scapose. Perennials; (cespitose, sometimes forming mats); caudex branched (with persistent leaf bases, branches creeping, terminating in scapes or sterile rosettes); scapose.
Stems

unbranched, (0.3–)0.5–1.7(–2.8) dm, pubescent throughout, trichomes simple and 2-rayed, 0.3–0.8 mm, with 3–5-rayed ones, 0.1–0.3 mm.

unbranched, (0.3–)0.4–1.6(–1.9) dm, pubescent, trichomes sessile, pectinate, 0.1–0.4 mm, (parallel to long axis of stem, sometimes with irregularly 2–4-rayed ones, 0.2–0.6 mm).

Basal leaves

rosulate; petiolate;

petiole base (not thickened), ciliate, (trichomes simple, 0.3–1 mm);

blade oblanceolate to obovate or lanceolate to oblong, 0.8–3(–4.5) cm × 2.5–6(–9) mm, margins entire, surfaces abaxially pubescent with stalked, 2–4-rayed trichomes, 0.1–0.5 mm, with simple ones (midvein obscure, not thickened), adaxially glabrous or sparsely pubescent with simple and stalked, 2–4-rayed trichomes.

rosulate;

subsessile;

blade narrowly oblanceolate to linear, (0.4–)0.6–1.3(–2) cm × 0.9–2.2 mm, margins entire (not ciliate), surfaces pubescent with subsessile or sessile, pectinate trichomes, 0.2–0.5 mm.

Cauline leaves

0.

0.

Racemes

6–18-flowered, ebracteate, considerably elongated in fruit;

rachis not flexuous, pubescent as stem.

5–22-flowered, ebracteate, considerably elongated in fruit;

rachis not flexuous, pubescent as stem.

Flowers

sepals (purplish tinged), narrowly ovate, 2.5–3 mm, pubescent, (trichomes simple and fewer, stalked, 2-rayed);

petals bright yellow, narrowly obovate, 3.5–5 × 1.7–2.5 mm;

anthers ovate, 0.3–0.4 mm.

sepals broadly ovate, 2–3.2 mm, pubescent, (trichomes pectinate);

petals yellow, obovate to spatulate, 4–6.5 × 1.5–2.5 mm;

anthers ovate 0.4–0.5 mm.

Fruiting pedicels

ascending to divaricate-ascending, straight or, sometimes, slightly curved upwards, 4–14(–30) mm, pubescent, trichomes simple and 2–4-rayed.

divaricate-ascending, straight, (5–)7–14 mm, sparsely pubescent, trichomes pectinate.

Fruits

elliptic, plane, flattened, 6–10 × 2–3 mm;

valves glabrous or glabrescent, trichomes simple, (not confined to replum);

ovules 12–24 per ovary;

style 0.2–0.3 mm (stigma about as wide as style).

ovoid, plane, slightly inflated basally, 4–6(–7) × 2–3 mm;

valves pubescent, trichomes usually sessile, pectinate, 0.2–0.5 mm, rarely with simple ones;

ovules 4–8 per ovary;

style 0.5–1.5 mm.

Seeds

(pale brown), ovoid, 0.9–1.3 × 0.6–0.9 mm.

oblong, 1.1–1.5 × 0.6–0.8 mm.

2n

= 80.

= 22.

Draba alpina

Draba pectinipila

Phenology Flowering Jun–Aug. Flowering Apr–Jun.
Habitat Moist tundra and ridges, sand and gravel flats or beaches Rocky slopes in sagebrush scrub and pinyon-juniper woodlands
Elevation 0-1000 m (0-3300 ft) 1700-2400 m (5600-7900 ft)
Distribution
from FNA
LB; MB; NU; ON; QC; Greenland; Europe (Finland, Norway, Russia, Sweden)
[WildflowerSearch map]
[BONAP county map]
from FNA
CO; MT; UT; WY
[BONAP county map]
Discussion

The synonymy above includes two North American names overlooked by C. L. Hitchcock (1941) and R. C. Rollins (1993). Draba alpina was broadly delimited by O. E. Schulz (1927) and included 17 varieties, some of which (e.g., corymbosa, oxycarpa, pilosa) are recognized herein as distinct species. The name D. alpina was so misapplied that it was used for any circumpolar or alpine, scapose, yellow-flowered, perennial Draba. Various chromosome numbers (e.g., 2n = 64, 80, 112, 120; S. I. Warwick and I. A. Al-Shehbaz 2006) have been reported for the species. As circumscribed here, it has the narrow distribution outlined above and includes plants with 2n = 80. Reports of the species from Alaska, Canadian Northwest Territories and Yukon, Siberia, eastern Asia, Russian Far East, and the Central Asian republics are either suspect or very unlikely. The entire D. alpina complex (including the above three species, D. glacialis Adams, D. macounii, etc.) is in need of critical molecular, cytological, and morphological study.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Draba pectinipila was treated as a synonym of D. oligosperma by G. A. Mulligan (1972), R. C. Rollins (1993), and N. H. Holmgren (2005b). The latter recognized D. juniperina as a distinct species but overlooked the fact that it is indistinguishable from the type collections of the earlier-published D. pectinipila. The species (including D. juniperina) differs significantly from D. oligosperma in both chromosome number and morphology. Draba pectinipila is easily distinguished by having fruit valves pubescent with pectinate trichomes, fruiting pedicels (5–)7–14 mm, petals 4–6.5 mm, ovules 4–8 per ovary, and styles 0.5–1.5 mm. By contrast, D. oligosperma has fruit valves glabrous or pubescent with simple or 2-rayed trichomes, fruiting pedicels (2–)3–10(–13) mm, petals 2.5–4 mm, ovules 6–12 per ovary, and styles 0.1–0.8(–1.1) mm. Draba pectinipila was previously known only from the type locality in northwestern Wyoming (Park County). Its range is now expanded to include that of D. juniperina in northwestern Colorado (Moffat County), northeastern Utah (Daggett and Uintah counties), and southwestern Wyoming (Sweetwater County). The record from Uintah County is based on Goodrich 22275 (NY).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 290. FNA vol. 7, p. 328.
Parent taxa Brassicaceae > tribe Arabideae > Draba Brassicaceae > tribe Arabideae > Draba
Sibling taxa
D. abajoensis, D. albertina, D. aleutica, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. cyclomorpha, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. helleriana, D. hitchcockii, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis
D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. cyclomorpha, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. helleriana, D. hitchcockii, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis
Synonyms D. alpina var. hydeana, D. alpina var. inflatisiliqua D. juniperina, D. oligosperma var. juniperina, D. oligosperma var. pectinipila
Name authority Linnaeus: Sp. Pl. 2: 642. (1753) Rollins: Rhodora 55: 231. (1953)
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