Draba alpina |
Draba paucifructa |
|
---|---|---|
alpine Draba |
Charleston Mountain Draba |
|
Habit | Perennials; (cespitose); caudex branched (covered with persistent leaves or leaf remains); scapose. | Perennials; caudex simple or branched (not fleshy, usually with some persistent leaf remains); sometimes scapose. |
Stems | unbranched, (0.3–)0.5–1.7(–2.8) dm, pubescent throughout, trichomes simple and 2-rayed, 0.3–0.8 mm, with 3–5-rayed ones, 0.1–0.3 mm. |
unbranched, (0.1–)0.3–0.9 dm, glabrous throughout or sparsely pubescent proximally, trichomes simple and 2–6-rayed, 0.1–0.5 mm. |
Basal leaves | rosulate; petiolate; petiole base (not thickened), ciliate, (trichomes simple, 0.3–1 mm); blade oblanceolate to obovate or lanceolate to oblong, 0.8–3(–4.5) cm × 2.5–6(–9) mm, margins entire, surfaces abaxially pubescent with stalked, 2–4-rayed trichomes, 0.1–0.5 mm, with simple ones (midvein obscure, not thickened), adaxially glabrous or sparsely pubescent with simple and stalked, 2–4-rayed trichomes. |
rosulate; petiolate; petiole (0–0.5 cm), ciliate throughout; blade oblanceolate to obovate, 0.4–1.7(–2.3) cm × 1.6–4(–6) mm, margins usually entire, rarely denticulate, (sparsely ciliate, trichomes simple, 0.3–0.8 mm), surfaces pubescent, abaxially with stalked, (2–)4(–6)-rayed trichomes, (0.05–)0.1–0.6 mm, adaxially with simple and fewer, stalked, 2–4-rayed ones. |
Cauline leaves | 0. |
0 or 1; sessile; blade ovate to lanceolate, margins entire, surfaces pubescent as basal. |
Racemes | 6–18-flowered, ebracteate, considerably elongated in fruit; rachis not flexuous, pubescent as stem. |
(2–)4–9(–12)-flowered, ebracteate, elongated in fruit; rachis slightly flexuous, glabrous. |
Flowers | sepals (purplish tinged), narrowly ovate, 2.5–3 mm, pubescent, (trichomes simple and fewer, stalked, 2-rayed); petals bright yellow, narrowly obovate, 3.5–5 × 1.7–2.5 mm; anthers ovate, 0.3–0.4 mm. |
sepals ovate, 1.2–1.7 mm, pubescent, (trichomes simple and short-stalked, 2- or 3-rayed); petals yellowish (quickly fading white), spatulate, 1.8–2.2 × 0.6–1 mm; anthers ovate, 0.25–0.3 mm. |
Fruiting pedicels | ascending to divaricate-ascending, straight or, sometimes, slightly curved upwards, 4–14(–30) mm, pubescent, trichomes simple and 2–4-rayed. |
divaricate-ascending, straight, 2–5(–8) mm, glabrous. |
Fruits | elliptic, plane, flattened, 6–10 × 2–3 mm; valves glabrous or glabrescent, trichomes simple, (not confined to replum); ovules 12–24 per ovary; style 0.2–0.3 mm (stigma about as wide as style). |
elliptic to linear lanceolate, plane, flattened, 5–10(–12) × 1.7–2(–2.5) mm; valves glabrous; ovules 20–30 per ovary; style 0.08–0.2 mm. |
Seeds | (pale brown), ovoid, 0.9–1.3 × 0.6–0.9 mm. |
oblong, 0.9–1.2 × 0.5–0.8 mm. |
2n | = 80. |
= 40. |
Draba alpina |
Draba paucifructa |
|
Phenology | Flowering Jun–Aug. | Flowering Jun–Jul. |
Habitat | Moist tundra and ridges, sand and gravel flats or beaches | Moist, shaded slopes among limestone and dolomite rocks |
Elevation | 0-1000 m (0-3300 ft) | 2600-3500 m (8500-11500 ft) |
Distribution |
LB; MB; NU; ON; QC; Greenland; Europe (Finland, Norway, Russia, Sweden)
|
NV |
Discussion | The synonymy above includes two North American names overlooked by C. L. Hitchcock (1941) and R. C. Rollins (1993). Draba alpina was broadly delimited by O. E. Schulz (1927) and included 17 varieties, some of which (e.g., corymbosa, oxycarpa, pilosa) are recognized herein as distinct species. The name D. alpina was so misapplied that it was used for any circumpolar or alpine, scapose, yellow-flowered, perennial Draba. Various chromosome numbers (e.g., 2n = 64, 80, 112, 120; S. I. Warwick and I. A. Al-Shehbaz 2006) have been reported for the species. As circumscribed here, it has the narrow distribution outlined above and includes plants with 2n = 80. Reports of the species from Alaska, Canadian Northwest Territories and Yukon, Siberia, eastern Asia, Russian Far East, and the Central Asian republics are either suspect or very unlikely. The entire D. alpina complex (including the above three species, D. glacialis Adams, D. macounii, etc.) is in need of critical molecular, cytological, and morphological study. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Draba paucifructa is known from the Charleston Mountains in Clark County. M. D. Windham (2004) suggested that it is an allopolyploid resulting from hybridization between D. albertina and D. lonchocarpa, and he discussed the morphological features that distinguish this species from its putative parents. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 290. | FNA vol. 7, p. 327. |
Parent taxa | Brassicaceae > tribe Arabideae > Draba | Brassicaceae > tribe Arabideae > Draba |
Sibling taxa | ||
Synonyms | D. alpina var. hydeana, D. alpina var. inflatisiliqua | |
Name authority | Linnaeus: Sp. Pl. 2: 642. (1753) | Clokey & C. L. Hitchcock: Madroño 5: 127. (1939) |
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