Draba alpina |
Draba norvegica |
|
---|---|---|
alpine Draba |
Norwegian Draba |
|
Habit | Perennials; (cespitose); caudex branched (covered with persistent leaves or leaf remains); scapose. | Perennials; (cespitose); caudex branched (sometimes with persistent leaf remains); often scapose. |
Stems | unbranched, (0.3–)0.5–1.7(–2.8) dm, pubescent throughout, trichomes simple and 2-rayed, 0.3–0.8 mm, with 3–5-rayed ones, 0.1–0.3 mm. |
branched or unbranched, (0.2–)0.4–1.4(–2) dm, pubescent throughout, trichomes simple and 2–4-rayed, 0.2–0.5 mm. |
Basal leaves | rosulate; petiolate; petiole base (not thickened), ciliate, (trichomes simple, 0.3–1 mm); blade oblanceolate to obovate or lanceolate to oblong, 0.8–3(–4.5) cm × 2.5–6(–9) mm, margins entire, surfaces abaxially pubescent with stalked, 2–4-rayed trichomes, 0.1–0.5 mm, with simple ones (midvein obscure, not thickened), adaxially glabrous or sparsely pubescent with simple and stalked, 2–4-rayed trichomes. |
rosulate; petiolate; petiole (0–0.5 cm), ciliate throughout; blade oblanceolate to narrowly obovate or narrowly lanceolate, 0.4–2.1 cm × 1.5–7 mm, margins with 1–3 teeth on each side, (ciliate, trichomes simple and 2-rayed, 0.2–0.8 mm), surfaces pubescent abaxially with stalked, (2–)4(–6)-rayed trichomes, 0.1–0.4 mm, adaxially glabrous or pubescent, with simple and stalked, 2-rayed trichomes, 0.1–0.5 mm. |
Cauline leaves | 0. |
0–3(–5); sessile; blade ovate to oblong, margins often dentate, (ciliate). |
Racemes | 6–18-flowered, ebracteate, considerably elongated in fruit; rachis not flexuous, pubescent as stem. |
5–23-flowered, ebracteate or proximalmost 1(–3) flowers bracteate, considerably elongated in fruit; rachis slightly or not flexuous, pubescent as stem. |
Flowers | sepals (purplish tinged), narrowly ovate, 2.5–3 mm, pubescent, (trichomes simple and fewer, stalked, 2-rayed); petals bright yellow, narrowly obovate, 3.5–5 × 1.7–2.5 mm; anthers ovate, 0.3–0.4 mm. |
sepals ovate, 1.7–2.5 mm, pubescent, (trichomes simple and short-stalked, 2–4-rayed); petals white, spatulate, 2.5–4 × 1.7–2.5 mm; anthers ovate, 0.3–0.4 mm. |
Fruiting pedicels | ascending to divaricate-ascending, straight or, sometimes, slightly curved upwards, 4–14(–30) mm, pubescent, trichomes simple and 2–4-rayed. |
suberect to ascending (often appressed to rachis), straight, (1.5–)2.5–5(–9) mm, pubescent, trichomes simple and 2–4-rayed. |
Fruits | elliptic, plane, flattened, 6–10 × 2–3 mm; valves glabrous or glabrescent, trichomes simple, (not confined to replum); ovules 12–24 per ovary; style 0.2–0.3 mm (stigma about as wide as style). |
(sometimes subappressed to rachis), oblong to lanceolate-elliptic, plane, flattened, 4–8 × 2–3 mm; valves glabrous or pubescent, trichomes simple and short-stalked, 2-rayed, 0.05–0.25 mm; ovules 18–26 per ovary; style (0.01–)0.1–0.4(–0.5) mm, (glabrous, stigma about as wide as style). |
Seeds | (pale brown), ovoid, 0.9–1.3 × 0.6–0.9 mm. |
oblong, 0.9–1.2 × 0.5–0.7 mm. |
2n | = 80. |
= 48. |
Draba alpina |
Draba norvegica |
|
Phenology | Flowering Jun–Aug. | Flowering Jun–Aug. |
Habitat | Moist tundra and ridges, sand and gravel flats or beaches | Rock outcrops and sea cliffs, talus, gravelly and sandy terraces, moist bluffs, turfy limestone shores |
Elevation | 0-1000 m (0-3300 ft) | 0-500 m (0-1600 ft) |
Distribution |
LB; MB; NU; ON; QC; Greenland; Europe (Finland, Norway, Russia, Sweden)
|
NL; NS; NT; QC; Greenland; Europe (Finland, Norway, n Russia, Scotland, Sweden); Atlantic Islands (Iceland) |
Discussion | The synonymy above includes two North American names overlooked by C. L. Hitchcock (1941) and R. C. Rollins (1993). Draba alpina was broadly delimited by O. E. Schulz (1927) and included 17 varieties, some of which (e.g., corymbosa, oxycarpa, pilosa) are recognized herein as distinct species. The name D. alpina was so misapplied that it was used for any circumpolar or alpine, scapose, yellow-flowered, perennial Draba. Various chromosome numbers (e.g., 2n = 64, 80, 112, 120; S. I. Warwick and I. A. Al-Shehbaz 2006) have been reported for the species. As circumscribed here, it has the narrow distribution outlined above and includes plants with 2n = 80. Reports of the species from Alaska, Canadian Northwest Territories and Yukon, Siberia, eastern Asia, Russian Far East, and the Central Asian republics are either suspect or very unlikely. The entire D. alpina complex (including the above three species, D. glacialis Adams, D. macounii, etc.) is in need of critical molecular, cytological, and morphological study. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Draba norvegica is a highly variable hexaploid taxon and, as delimited herein, it probably represents two taxa with the same distribution and same chromosome number. It is related to hexaploid D. arctogena, from which it is distinguished by characteristics listed under 8. D. arctogena. Draba norvegica was divided by O. E. Schulz (1927) into eight varieties, whereas R. C. Rollins (1993) recognized just two. In the latter treatment, var. clivicola is distinguished by having predominantly branched, appressed trichomes on stems proximally, lanceolate, glabrous fruits, and slender, erect-appressed pedicels. By contrast, var. norvegica is said to have predominantly simple, spreading trichomes on stems proximally, narrowly ovate to elliptical, glabrous or pubescent fruits, and stout, divaricately ascending pedicels. These differences are extremes of a continuum, and we agree with G. A. Mulligan (1970) in reducing D. clivicola to synonymy of D. norvegica. Rollins indicated that D. norvegica grows in Minnesota, but we have not seen any material from that state. Draba norvegica is found in Europe in arctic and subarctic areas. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 290. | FNA vol. 7, p. 322. |
Parent taxa | Brassicaceae > tribe Arabideae > Draba | Brassicaceae > tribe Arabideae > Draba |
Sibling taxa | ||
Synonyms | D. alpina var. hydeana, D. alpina var. inflatisiliqua | D. clivicola, D. hirta var. norvegica, D. norvegica var. clivicola, D. norvegica var. hebecarpa, D. rupestris, D. scandinavica var. hebecarpa, D. trichella |
Name authority | Linnaeus: Sp. Pl. 2: 642. (1753) | Gunnerus: Fl. Norveg. 2: 106. (1772) |
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