Draba alpina |
Draba graminea |
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alpine Draba |
Rocky Mountain Draba |
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Habit | Perennials; (cespitose); caudex branched (covered with persistent leaves or leaf remains); scapose. | Perennials; (cespitose); caudex simple or branched (with persistent leaf bases); not scapose. |
Stems | unbranched, (0.3–)0.5–1.7(–2.8) dm, pubescent throughout, trichomes simple and 2-rayed, 0.3–0.8 mm, with 3–5-rayed ones, 0.1–0.3 mm. |
unbranched, 0.1–0.5(–0.8) dm, usually pubescent throughout, rarely glabrous, trichomes simple or subsessile, 2-rayed or spurred, (crisped), 0.1–0.5 mm. |
Basal leaves | rosulate; petiolate; petiole base (not thickened), ciliate, (trichomes simple, 0.3–1 mm); blade oblanceolate to obovate or lanceolate to oblong, 0.8–3(–4.5) cm × 2.5–6(–9) mm, margins entire, surfaces abaxially pubescent with stalked, 2–4-rayed trichomes, 0.1–0.5 mm, with simple ones (midvein obscure, not thickened), adaxially glabrous or sparsely pubescent with simple and stalked, 2–4-rayed trichomes. |
rosulate; petiolate; petiole ciliate, (trichomes usually straight and simple, rarely also 2-rayed, 0.2–0.6(–0.8) mm); blade linear to linear-oblanceolate, (0.5–)1–4 cm × 0.3–2(–3) mm, margins entire (pubescent as petiole), surfaces glabrous. |
Cauline leaves | 0. |
(1–)3–9(–12) (as bracts); sessile; blade linear to oblanceolate or lanceolate, margins entire, (ciliate proximally, similar to basal). |
Racemes | 6–18-flowered, ebracteate, considerably elongated in fruit; rachis not flexuous, pubescent as stem. |
3–15-flowered, bracteate, elongated in fruit; rachis not flexuous, usually pubescent as stem, rarely glabrous. |
Flowers | sepals (purplish tinged), narrowly ovate, 2.5–3 mm, pubescent, (trichomes simple and fewer, stalked, 2-rayed); petals bright yellow, narrowly obovate, 3.5–5 × 1.7–2.5 mm; anthers ovate, 0.3–0.4 mm. |
sepals ovate, 1.5–2.5 mm, glabrous or pubescent, (trichomes simple and short-stalked, 2-rayed); petals yellow, spatulate to obovate, 3–5 × 1.5–3 mm; anthers ovate, 0.3–0.5 mm. |
Fruiting pedicels | ascending to divaricate-ascending, straight or, sometimes, slightly curved upwards, 4–14(–30) mm, pubescent, trichomes simple and 2–4-rayed. |
divaricate-ascending, usually straight, rarely curved upward, 3–10(–15) mm, usually pubescent as stem, rarely glabrous. |
Fruits | elliptic, plane, flattened, 6–10 × 2–3 mm; valves glabrous or glabrescent, trichomes simple, (not confined to replum); ovules 12–24 per ovary; style 0.2–0.3 mm (stigma about as wide as style). |
ovate-elliptic to lanceolate, slightly twisted or plane, flattened, 5–11 × 2.5–5 mm; valves glabrous; ovules 8–16 per ovary; style 0.2–0.7 mm. |
Seeds | (pale brown), ovoid, 0.9–1.3 × 0.6–0.9 mm. |
ovoid, 1.2–1.5 × 0.7–1 mm. |
2n | = 80. |
= 18. |
Draba alpina |
Draba graminea |
|
Phenology | Flowering Jun–Aug. | Flowering Jul–Sep. |
Habitat | Moist tundra and ridges, sand and gravel flats or beaches | Rocky areas and ridges, alpine tundra, gravel bars in streams |
Elevation | 0-1000 m (0-3300 ft) | 3000-4100 m (9800-13500 ft) |
Distribution |
LB; MB; NU; ON; QC; Greenland; Europe (Finland, Norway, Russia, Sweden)
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CO
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Discussion | The synonymy above includes two North American names overlooked by C. L. Hitchcock (1941) and R. C. Rollins (1993). Draba alpina was broadly delimited by O. E. Schulz (1927) and included 17 varieties, some of which (e.g., corymbosa, oxycarpa, pilosa) are recognized herein as distinct species. The name D. alpina was so misapplied that it was used for any circumpolar or alpine, scapose, yellow-flowered, perennial Draba. Various chromosome numbers (e.g., 2n = 64, 80, 112, 120; S. I. Warwick and I. A. Al-Shehbaz 2006) have been reported for the species. As circumscribed here, it has the narrow distribution outlined above and includes plants with 2n = 80. Reports of the species from Alaska, Canadian Northwest Territories and Yukon, Siberia, eastern Asia, Russian Far East, and the Central Asian republics are either suspect or very unlikely. The entire D. alpina complex (including the above three species, D. glacialis Adams, D. macounii, etc.) is in need of critical molecular, cytological, and morphological study. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Draba graminea is occasionally confused with D. crassa, which occupies similar habitats and elevations. It is easily distinguished from the latter by having narrower [0.3–2(–3) versus 2.5–8(–10) mm wide] basal leaves and bracteate (versus ebracteate) racemes. It is known to us only from Hinsdale, La Plata, Ouray, San Juan, and San Miguel counties. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 290. | FNA vol. 7, p. 309. |
Parent taxa | Brassicaceae > tribe Arabideae > Draba | Brassicaceae > tribe Arabideae > Draba |
Sibling taxa | ||
Synonyms | D. alpina var. hydeana, D. alpina var. inflatisiliqua | Braya graminea, D. chrysantha var. graminea, D. chrysantha var. hirticaulis |
Name authority | Linnaeus: Sp. Pl. 2: 642. (1753) | Greene: Pl. Baker. 3: 5. (1901) |
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