Flora of North America species comparison

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Draba alpina

Draba cusickii

alpine Draba

Cusick's Draba, Steens Mountain whitlow-grass
Habit Perennials; (cespitose); caudex branched (covered with persistent leaves or leaf remains); scapose. Perennials; (loosely cespitose); caudex branched (with few, persistent leaf remains); scapose.
Stems

unbranched, (0.3–)0.5–1.7(–2.8) dm, pubescent throughout, trichomes simple and 2-rayed, 0.3–0.8 mm, with 3–5-rayed ones, 0.1–0.3 mm.

unbranched, 0.3–1.2 dm, pubescent throughout, trichomes (2–)4-rayed, 0.05–0.5(–0.6) mm.
Basal leaves

rosulate; petiolate;

petiole base (not thickened), ciliate, (trichomes simple, 0.3–1 mm);

blade oblanceolate to obovate or lanceolate to oblong, 0.8–3(–4.5) cm × 2.5–6(–9) mm, margins entire, surfaces abaxially pubescent with stalked, 2–4-rayed trichomes, 0.1–0.5 mm, with simple ones (midvein obscure, not thickened), adaxially glabrous or sparsely pubescent with simple and stalked, 2–4-rayed trichomes.

rosulate; shortly petiolate;

petiole base ciliate (margin not ciliate, trichomes 2-rayed, 0.3–0.6 mm);

blade oblanceolate to obovate, 0.4–1.2 cm × 2–5 mm, margins entire or, rarely, denticulate, surfaces pubescent with stalked, (2–)4-rayed trichomes, 0.15–0.4 mm.
Cauline leaves

0.

0.
Racemes

6–18-flowered, ebracteate, considerably elongated in fruit;

rachis not flexuous, pubescent as stem.

10–20(–25)-flowered, ebracteate, considerably elongated in fruit;

rachis not flexuous, pubescent as stems proximally.
Flowers

sepals (purplish tinged), narrowly ovate, 2.5–3 mm, pubescent, (trichomes simple and fewer, stalked, 2-rayed);

petals bright yellow, narrowly obovate, 3.5–5 × 1.7–2.5 mm;

anthers ovate, 0.3–0.4 mm.

sepals broadly ovate, 2–2.5 mm, pubescent, (trichomes short-stalked, 3- or 4-rayed);

petals yellow, obovate to oblanceolate, 3.7–6 × 1.5–2.5 mm;

anthers ovate, 0.5–0.6 mm.
Fruiting pedicels

ascending to divaricate-ascending, straight or, sometimes, slightly curved upwards, 4–14(–30) mm, pubescent, trichomes simple and 2–4-rayed.

divaricate-ascending, straight (not expanded basally), 4–9 mm, pubescent, trichomes (2–)4-rayed, (0.05–0.25 mm).
Fruits

elliptic, plane, flattened, 6–10 × 2–3 mm;

valves glabrous or glabrescent, trichomes simple, (not confined to replum);

ovules 12–24 per ovary;

style 0.2–0.3 mm (stigma about as wide as style).

lanceolate to broadly ovate, plane, flattened, 6–9 × 2–4.5 mm;

valves pubescent, trichomes short-stalked, (2–)4-rayed, 0.05–0.25 mm;

ovules 10–18 per ovary;

style 0.4 0.8(–1) mm.
Seeds

(pale brown), ovoid, 0.9–1.3 × 0.6–0.9 mm.

oblong, 1.2–1.5 × 0.7–0.8 mm.
2n

= 80.

= 26.

Draba alpina

Draba cusickii
Phenology Flowering Jun–Aug. Flowering Jun–Jul.
Habitat Moist tundra and ridges, sand and gravel flats or beaches Basaltic outcrops, talus, rocky soil
Elevation 0-1000 m (0-3300 ft) 2000-2700 m (6600-8900 ft)
Distribution
from FNA
LB; MB; NU; ON; QC; Greenland; Europe (Finland, Norway, Russia, Sweden)
[WildflowerSearch map]
[BONAP county map]
from FNA
OR
[WildflowerSearch map]
[BONAP county map]
Discussion

The synonymy above includes two North American names overlooked by C. L. Hitchcock (1941) and R. C. Rollins (1993). Draba alpina was broadly delimited by O. E. Schulz (1927) and included 17 varieties, some of which (e.g., corymbosa, oxycarpa, pilosa) are recognized herein as distinct species. The name D. alpina was so misapplied that it was used for any circumpolar or alpine, scapose, yellow-flowered, perennial Draba. Various chromosome numbers (e.g., 2n = 64, 80, 112, 120; S. I. Warwick and I. A. Al-Shehbaz 2006) have been reported for the species. As circumscribed here, it has the narrow distribution outlined above and includes plants with 2n = 80. Reports of the species from Alaska, Canadian Northwest Territories and Yukon, Siberia, eastern Asia, Russian Far East, and the Central Asian republics are either suspect or very unlikely. The entire D. alpina complex (including the above three species, D. glacialis Adams, D. macounii, etc.) is in need of critical molecular, cytological, and morphological study.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Draba cusickii was divided by R. C. Rollins (1993) into two varieties with significantly different morphologies and chromosome numbers. The Nevada endemic var. pedicellata (2n = 20) is here treated as a distinct species following M. D. Windham (2004). Draba cusickii has the same chromosome number as D. sobolifera but is readily distinguished from that species by having 4-rayed (versus simple or 2-rayed) fruit trichomes. Draba cusickii is known from Steens Mountain (Harney County).

(Discussion copyrighted by Flora of North America; reprinted with permission.)
Source FNA vol. 7, p. 290. FNA vol. 7, p. 304.
Parent taxa Brassicaceae > tribe Arabideae > Draba Brassicaceae > tribe Arabideae > Draba
Sibling taxa
D. abajoensis, D. albertina, D. aleutica, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. cyclomorpha, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. helleriana, D. hitchcockii, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis
D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cyclomorpha, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. helleriana, D. hitchcockii, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis
Synonyms D. alpina var. hydeana, D. alpina var. inflatisiliqua D. sphaeroides var. cusickii
Name authority Linnaeus: Sp. Pl. 2: 642. (1753) B. L. Robinson ex O. E. Schulz: in H. G. A. Engler, Pflanzenr. 89[IV,105]: 105. (1927)
Web links 
Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).



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