FNA: Draba alpina vs. Draba corrugata

	Draba alpina	Draba corrugata
	alpine Draba	Southern California Draba
Habit	Perennials; (cespitose); caudex branched (covered with persistent leaves or leaf remains); scapose.	Biennials or perennials; (short-lived); caudex simple (covered with persistent leaves); not scapose.
Stems	unbranched, (0.3–)0.5–1.7(–2.8) dm, pubescent throughout, trichomes simple and 2-rayed, 0.3–0.8 mm, with 3–5-rayed ones, 0.1–0.3 mm.	branched, (0.3–) 0.4–1.7(–2.5) dm, pubescent throughout, trichomes simple and long-stalked, 2-rayed, 0.4–1.4 mm, with smaller, 2–4-rayed ones, 0.1–0.6 mm, (simple ones usually fewer distally).
Basal leaves	rosulate; petiolate; petiole base (not thickened), ciliate, (trichomes simple, 0.3–1 mm); blade oblanceolate to obovate or lanceolate to oblong, 0.8–3(–4.5) cm × 2.5–6(–9) mm, margins entire, surfaces abaxially pubescent with stalked, 2–4-rayed trichomes, 0.1–0.5 mm, with simple ones (midvein obscure, not thickened), adaxially glabrous or sparsely pubescent with simple and stalked, 2–4-rayed trichomes.	(densely imbricate); rosulate; shortly petiolate; petiole base and margin ciliate, (trichomes simple, 0.6–2 mm); blade oblong to narrowly oblanceolate, (0.5–)1–2.2(–4.5) cm × 2–5 mm, margins entire, surfaces densely pubescent, abaxially with stalked, 2–4-rayed trichomes, 0.4–1.2 mm, (simple trichomes often along midvein), adaxially with mostly simple and long-stalked, 2-rayed trichomes, 0.6–1.3 mm, sometimes with 3- or 4-rayed ones.
Cauline leaves	0.	(4–)6–10(–13); sessile; blade oblong to ovate, margins entire, surfaces pubescent as basal.
Racemes	6–18-flowered, ebracteate, considerably elongated in fruit; rachis not flexuous, pubescent as stem.	(10–)18–55(–67)-flowered, ebracteate or proximalmost flowers bracteate, slightly or considerably elongated in fruit; rachis not flexuous, pubescent as stem.
Flowers	sepals (purplish tinged), narrowly ovate, 2.5–3 mm, pubescent, (trichomes simple and fewer, stalked, 2-rayed); petals bright yellow, narrowly obovate, 3.5–5 × 1.7–2.5 mm; anthers ovate, 0.3–0.4 mm.	sepals oblong, 2–2.7 mm, pubescent, (trichomes short-stalked, 2–4-rayed); petals yellow, linear, 2–3.5 × 0.2–0.5 mm; anthers oblong, 0.6–0.8 mm (exserted).
Fruiting pedicels	ascending to divaricate-ascending, straight or, sometimes, slightly curved upwards, 4–14(–30) mm, pubescent, trichomes simple and 2–4-rayed.	divaricate to ascending, straight, 2–6(–8) mm, pubescent as stem.
Fruits	elliptic, plane, flattened, 6–10 × 2–3 mm; valves glabrous or glabrescent, trichomes simple, (not confined to replum); ovules 12–24 per ovary; style 0.2–0.3 mm (stigma about as wide as style).	elliptic to oblong or linear- to oblong-elliptic, slightly twisted or plane, flattened, 5–13(–17) × 2–3(–4) mm; valves pubescent, trichomes short-stalked, cruciform, 0.1–0.5 mm, (sometimes with 2- or 3-rayed ones); ovules 16–28 per ovary; style 1.4–3.4 mm.
Seeds	(pale brown), ovoid, 0.9–1.3 × 0.6–0.9 mm.	oblong, 1–1.2 × 0.6–0.8 mm.
2n	= 80.

	Draba alpina	Draba corrugata
Phenology	Flowering Jun–Aug.	Flowering Jun–Jul.
Habitat	Moist tundra and ridges, sand and gravel flats or beaches	Alpine fellfields, talus, open pine woodlands
Elevation	0-1000 m (0-3300 ft)	2000-3500 m (6600-11500 ft)
Distribution	LB; MB; NU; ON; QC; Greenland; Europe (Finland, Norway, Russia, Sweden) [WildflowerSearch map] [BONAP county map]	CA [WildflowerSearch map] [BONAP county map]
Discussion	The synonymy above includes two North American names overlooked by C. L. Hitchcock (1941) and R. C. Rollins (1993). Draba alpina was broadly delimited by O. E. Schulz (1927) and included 17 varieties, some of which (e.g., corymbosa, oxycarpa, pilosa) are recognized herein as distinct species. The name D. alpina was so misapplied that it was used for any circumpolar or alpine, scapose, yellow-flowered, perennial Draba. Various chromosome numbers (e.g., 2n = 64, 80, 112, 120; S. I. Warwick and I. A. Al-Shehbaz 2006) have been reported for the species. As circumscribed here, it has the narrow distribution outlined above and includes plants with 2n = 80. Reports of the species from Alaska, Canadian Northwest Territories and Yukon, Siberia, eastern Asia, Russian Far East, and the Central Asian republics are either suspect or very unlikely. The entire D. alpina complex (including the above three species, D. glacialis Adams, D. macounii, etc.) is in need of critical molecular, cytological, and morphological study. (Discussion copyrighted by Flora of North America; reprinted with permission.)	Of conservation concern. Draba corrugata was broadly circumscribed by R. C. Rollins (1993) to include three varieties that we treat as distinct species. For a discussion of species limits and distinguishing features, see I. A. Al-Shehbaz and M. D. Windham (2007). Draba corrugata, in the strict sense, is known from the San Antonio, San Bernardino, and San Gabriel mountains in Los Angeles and San Bernardino counties. (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 7, p. 290.	FNA vol. 7, p. 301.
Parent taxa	Brassicaceae > tribe Arabideae > Draba	Brassicaceae > tribe Arabideae > Draba
Sibling taxa	D. abajoensis, D. albertina, D. aleutica, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. cyclomorpha, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. helleriana, D. hitchcockii, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis	D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. cyclomorpha, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. helleriana, D. hitchcockii, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis
Synonyms	D. alpina var. hydeana, D. alpina var. inflatisiliqua	D. vestita
Name authority	Linnaeus: Sp. Pl. 2: 642. (1753)	S. Watson: in W. H. Brewer et al., Bot. California 2: 430. (1880)
Web links	WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	Local floras: CA Local Web sites: CalFlora, CalPhotos WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Draba corrugata

alpine Draba

Southern California Draba

Habit

Perennials; (cespitose); caudex branched (covered with persistent leaves or leaf remains); scapose.

Biennials or perennials; (short-lived); caudex simple (covered with persistent leaves); not scapose.

Stems

unbranched, (0.3–)0.5–1.7(–2.8) dm, pubescent throughout, trichomes simple and 2-rayed, 0.3–0.8 mm, with 3–5-rayed ones, 0.1–0.3 mm.

branched, (0.3–) 0.4–1.7(–2.5) dm, pubescent throughout, trichomes simple and long-stalked, 2-rayed, 0.4–1.4 mm, with smaller, 2–4-rayed ones, 0.1–0.6 mm, (simple ones usually fewer distally).

Basal leaves

rosulate; petiolate;

petiole base (not thickened), ciliate, (trichomes simple, 0.3–1 mm);

blade oblanceolate to obovate or lanceolate to oblong, 0.8–3(–4.5) cm × 2.5–6(–9) mm, margins entire, surfaces abaxially pubescent with stalked, 2–4-rayed trichomes, 0.1–0.5 mm, with simple ones (midvein obscure, not thickened), adaxially glabrous or sparsely pubescent with simple and stalked, 2–4-rayed trichomes.

(densely imbricate); rosulate; shortly petiolate;

petiole base and margin ciliate, (trichomes simple, 0.6–2 mm);

blade oblong to narrowly oblanceolate, (0.5–)1–2.2(–4.5) cm × 2–5 mm, margins entire, surfaces densely pubescent, abaxially with stalked, 2–4-rayed trichomes, 0.4–1.2 mm, (simple trichomes often along midvein), adaxially with mostly simple and long-stalked, 2-rayed trichomes, 0.6–1.3 mm, sometimes with 3- or 4-rayed ones.

Cauline leaves

(4–)6–10(–13);

sessile;

blade oblong to ovate, margins entire, surfaces pubescent as basal.

Racemes

6–18-flowered, ebracteate, considerably elongated in fruit;

rachis not flexuous, pubescent as stem.

(10–)18–55(–67)-flowered, ebracteate or proximalmost flowers bracteate, slightly or considerably elongated in fruit;

rachis not flexuous, pubescent as stem.

Flowers

sepals (purplish tinged), narrowly ovate, 2.5–3 mm, pubescent, (trichomes simple and fewer, stalked, 2-rayed);

petals bright yellow, narrowly obovate, 3.5–5 × 1.7–2.5 mm;

anthers ovate, 0.3–0.4 mm.

sepals oblong, 2–2.7 mm, pubescent, (trichomes short-stalked, 2–4-rayed);

petals yellow, linear, 2–3.5 × 0.2–0.5 mm;

anthers oblong, 0.6–0.8 mm (exserted).

Fruiting pedicels

ascending to divaricate-ascending, straight or, sometimes, slightly curved upwards, 4–14(–30) mm, pubescent, trichomes simple and 2–4-rayed.

divaricate to ascending, straight, 2–6(–8) mm, pubescent as stem.

Fruits

elliptic, plane, flattened, 6–10 × 2–3 mm;

valves glabrous or glabrescent, trichomes simple, (not confined to replum);

ovules 12–24 per ovary;

style 0.2–0.3 mm (stigma about as wide as style).

elliptic to oblong or linear- to oblong-elliptic, slightly twisted or plane, flattened, 5–13(–17) × 2–3(–4) mm;

valves pubescent, trichomes short-stalked, cruciform, 0.1–0.5 mm, (sometimes with 2- or 3-rayed ones);

ovules 16–28 per ovary;

style 1.4–3.4 mm.

Seeds

(pale brown), ovoid, 0.9–1.3 × 0.6–0.9 mm.

oblong, 1–1.2 × 0.6–0.8 mm.

= 80.

Draba alpina

Draba corrugata

Phenology

Flowering Jun–Aug.

Flowering Jun–Jul.

Habitat

Moist tundra and ridges, sand and gravel flats or beaches

Alpine fellfields, talus, open pine woodlands

Elevation

0-1000 m (0-3300 ft)

2000-3500 m (6600-11500 ft)

Distribution

LB; MB; NU; ON; QC; Greenland; Europe (Finland, Norway, Russia, Sweden)

[WildflowerSearch map]

[BONAP county map]

[WildflowerSearch map]

[BONAP county map]

Discussion

The synonymy above includes two North American names overlooked by C. L. Hitchcock (1941) and R. C. Rollins (1993). Draba alpina was broadly delimited by O. E. Schulz (1927) and included 17 varieties, some of which (e.g., corymbosa, oxycarpa, pilosa) are recognized herein as distinct species. The name D. alpina was so misapplied that it was used for any circumpolar or alpine, scapose, yellow-flowered, perennial Draba. Various chromosome numbers (e.g., 2n = 64, 80, 112, 120; S. I. Warwick and I. A. Al-Shehbaz 2006) have been reported for the species. As circumscribed here, it has the narrow distribution outlined above and includes plants with 2n = 80. Reports of the species from Alaska, Canadian Northwest Territories and Yukon, Siberia, eastern Asia, Russian Far East, and the Central Asian republics are either suspect or very unlikely. The entire D. alpina complex (including the above three species, D. glacialis Adams, D. macounii, etc.) is in need of critical molecular, cytological, and morphological study.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Draba corrugata was broadly circumscribed by R. C. Rollins (1993) to include three varieties that we treat as distinct species. For a discussion of species limits and distinguishing features, see I. A. Al-Shehbaz and M. D. Windham (2007). Draba corrugata, in the strict sense, is known from the San Antonio, San Bernardino, and San Gabriel mountains in Los Angeles and San Bernardino counties.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 7, p. 290.

FNA vol. 7, p. 301.

Parent taxa

Brassicaceae > tribe Arabideae > Draba

Sibling taxa

D. abajoensis, D. albertina, D. aleutica, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. cyclomorpha, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. helleriana, D. hitchcockii, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis

D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. cyclomorpha, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. helleriana, D. hitchcockii, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis

Synonyms

D. alpina var. hydeana, D. alpina var. inflatisiliqua

D. vestita

Name authority

Linnaeus: Sp. Pl. 2: 642. (1753)

S. Watson: in W. H. Brewer et al., Bot. California 2: 430. (1880)

Web links

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

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Draba alpina

Draba corrugata

Draba alpina

Draba corrugata