Draba alpina |
Draba asterophora |
|
---|---|---|
alpine Draba |
Lake Tahoe Draba, Tahoe Draba |
|
Habit | Perennials; (cespitose); caudex branched (covered with persistent leaves or leaf remains); scapose. | Perennials; (loosely cespitose); caudex branched (somewhat surculose, with persistent leaf bases, branches sometimes terminating in sterile rosettes); scapose. |
Stems | unbranched, (0.3–)0.5–1.7(–2.8) dm, pubescent throughout, trichomes simple and 2-rayed, 0.3–0.8 mm, with 3–5-rayed ones, 0.1–0.3 mm. |
unbranched, 0.3–1.1 dm, glabrous throughout or sparsely pubescent proximally, trichomes (2–)4-rayed, 0.1–0.3 mm. |
Basal leaves | rosulate; petiolate; petiole base (not thickened), ciliate, (trichomes simple, 0.3–1 mm); blade oblanceolate to obovate or lanceolate to oblong, 0.8–3(–4.5) cm × 2.5–6(–9) mm, margins entire, surfaces abaxially pubescent with stalked, 2–4-rayed trichomes, 0.1–0.5 mm, with simple ones (midvein obscure, not thickened), adaxially glabrous or sparsely pubescent with simple and stalked, 2–4-rayed trichomes. |
rosulate; petiolate; petiole obsolete, margin rarely ciliate proximally; blade (somewhat fleshy), broadly obovate to suborbicular or spatulate, 0.4–1.4(–1.7) cm × (2–)3–6(–7) mm, margins entire, surfaces pubescent, trichomes stalked, cruciform, and 2-, 3-, or 5-rayed, 0.2–0.6 mm. |
Cauline leaves | 0. |
0 (or 1, as a bract). |
Racemes | 6–18-flowered, ebracteate, considerably elongated in fruit; rachis not flexuous, pubescent as stem. |
(5–)8–20(–27)-flowered, usually ebracteate, rarely proximalmost flowers bracteate, elongated in fruit; rachis not flexuous, glabrous. |
Flowers | sepals (purplish tinged), narrowly ovate, 2.5–3 mm, pubescent, (trichomes simple and fewer, stalked, 2-rayed); petals bright yellow, narrowly obovate, 3.5–5 × 1.7–2.5 mm; anthers ovate, 0.3–0.4 mm. |
sepals oblong, 3–4 mm, glabrous or sparsely pubescent, (trichomes subapical, short-stalked, 2–4-rayed); petals bright yellow, oblanceolate, 5–7 × 1.5–2.5 mm; anthers ovate, 0.5–0.6 mm. |
Fruiting pedicels | ascending to divaricate-ascending, straight or, sometimes, slightly curved upwards, 4–14(–30) mm, pubescent, trichomes simple and 2–4-rayed. |
horizontal to divaricate-ascending, straight or curved upward, 3–9 mm, glabrous. |
Fruits | elliptic, plane, flattened, 6–10 × 2–3 mm; valves glabrous or glabrescent, trichomes simple, (not confined to replum); ovules 12–24 per ovary; style 0.2–0.3 mm (stigma about as wide as style). |
lanceolate-ovate to broadly ovate or oblong, slightly twisted or plane, strongly flattened, 5–11(–14) × (3.5–)4–6 mm; valves glabrous or, rarely, puberulent, trichomes simple and short-stalked, 2- or 3-rayed, 0.05–0.3 mm; ovules 12–18 per ovary; style 0.2–1.6(–2) mm. |
Seeds | (pale brown), ovoid, 0.9–1.3 × 0.6–0.9 mm. |
(winged), ovate, 1.8–2.8 × 1.2–2 mm; (wing 0.5–1 mm wide). |
2n | = 80. |
= 40. |
Draba alpina |
Draba asterophora |
|
Phenology | Flowering Jun–Aug. | Flowering Jun–Aug. |
Habitat | Moist tundra and ridges, sand and gravel flats or beaches | Granitic rock outcrops, talus, gravelly soil |
Elevation | 0-1000 m (0-3300 ft) | 2600-3300 m (8500-10800 ft) |
Distribution |
LB; MB; NU; ON; QC; Greenland; Europe (Finland, Norway, Russia, Sweden)
|
CA; NV |
Discussion | The synonymy above includes two North American names overlooked by C. L. Hitchcock (1941) and R. C. Rollins (1993). Draba alpina was broadly delimited by O. E. Schulz (1927) and included 17 varieties, some of which (e.g., corymbosa, oxycarpa, pilosa) are recognized herein as distinct species. The name D. alpina was so misapplied that it was used for any circumpolar or alpine, scapose, yellow-flowered, perennial Draba. Various chromosome numbers (e.g., 2n = 64, 80, 112, 120; S. I. Warwick and I. A. Al-Shehbaz 2006) have been reported for the species. As circumscribed here, it has the narrow distribution outlined above and includes plants with 2n = 80. Reports of the species from Alaska, Canadian Northwest Territories and Yukon, Siberia, eastern Asia, Russian Far East, and the Central Asian republics are either suspect or very unlikely. The entire D. alpina complex (including the above three species, D. glacialis Adams, D. macounii, etc.) is in need of critical molecular, cytological, and morphological study. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
C. L. Hitchcock (1941) and R. C. Rollins (1993) divided Draba asterophora into two varieties based on minor differences in style and fruit lengths. These do not appear to define genetically discrete taxa. The species apparently is restricted to El Dorado County, California, and the Carson Range in Washoe County, Nevada. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 290. | FNA vol. 7, p. 294. |
Parent taxa | Brassicaceae > tribe Arabideae > Draba | Brassicaceae > tribe Arabideae > Draba |
Sibling taxa | ||
Synonyms | D. alpina var. hydeana, D. alpina var. inflatisiliqua | D. asterophora var. macrocarpa |
Name authority | Linnaeus: Sp. Pl. 2: 642. (1753) | Payson: Amer. J. Bot. 4: 263. (1917) |
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