|
alpine Draba
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arctic whitlowgrass
|
| Perennials; (cespitose); caudex branched (covered with persistent leaves or leaf remains); scapose. |
Perennials; (cespitose, not pulvinate); caudex simple or branched; rarely scapose. |
unbranched, (0.3–)0.5–1.7(–2.8) dm, pubescent throughout, trichomes simple and 2-rayed, 0.3–0.8 mm, with 3–5-rayed ones, 0.1–0.3 mm. |
unbranched, 0.3–1.7(–2.4) dm, pubescent, trichomes 4–10-rayed (rays sometimes branched), 0.1–0.3 mm, with fewer, simple ones, to 0.7 mm. |
rosulate; petiolate; petiole base (not thickened), ciliate, (trichomes simple, 0.3–1 mm); blade oblanceolate to obovate or lanceolate to oblong, 0.8–3(–4.5) cm × 2.5–6(–9) mm, margins entire, surfaces abaxially pubescent with stalked, 2–4-rayed trichomes, 0.1–0.5 mm, with simple ones (midvein obscure, not thickened), adaxially glabrous or sparsely pubescent with simple and stalked, 2–4-rayed trichomes. |
rosulate; petiolate; petiole base ciliate, (trichomes simple, 0.2–0.8 mm); blade oblanceolate to narrowly obovate, 0.4–2.3(–3) cm × 1.5–6.5 mm, margins usually entire, rarely with 1 tooth on each side, surfaces densely pubescent with minutely stalked, 8–12-rayed, stellate trichomes, 0.2–0.4 mm, sometimes with coarser, simple or 2-rayed ones, (midvein distinct abaxially). |
0. |
0 or 1(–3); sessile; blade ovate or oblong to lanceolate, margins entire, surfaces pubescent as basal. |
6–18-flowered, ebracteate, considerably elongated in fruit; rachis not flexuous, pubescent as stem. |
3–18(–25)-flowered, ebracteate, elongated in fruit; rachis not flexuous, pubescent as stem. |
sepals (purplish tinged), narrowly ovate, 2.5–3 mm, pubescent, (trichomes simple and fewer, stalked, 2-rayed); petals bright yellow, narrowly obovate, 3.5–5 × 1.7–2.5 mm; anthers ovate, 0.3–0.4 mm. |
sepals ovate, 2–2.5 mm, pubescent, (trichomes short-stalked, 2- or 3-rayed, and simple); petals white, spatulate to obovate, 3.5–6 × 1.8–2 mm; anthers ovate, 0.3–0.4 mm. |
ascending to divaricate-ascending, straight or, sometimes, slightly curved upwards, 4–14(–30) mm, pubescent, trichomes simple and 2–4-rayed. |
divaricate-ascending or ascending, straight, (1.5–)2–6 mm, pubescent, trichomes stalked, stellate, and, sometimes, simple. |
elliptic, plane, flattened, 6–10 × 2–3 mm; valves glabrous or glabrescent, trichomes simple, (not confined to replum); ovules 12–24 per ovary; style 0.2–0.3 mm (stigma about as wide as style). |
oblong to lanceolate, plane, slightly flattened, (5–)6–11 × 2–3 mm; valves pubescent, trichomes short-stalked, 2–5-rayed (some rays branched), 0.05–0.3 mm; ovules 20–32 per ovary; style 0.1–1 mm (stigma distinctly wider than style). |
(pale brown), ovoid, 0.9–1.3 × 0.6–0.9 mm. |
ovoid, 0.8–1.1 × (0.6–)0.7–0.8 mm. |
= 80. |
= 80. |
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| Flowering Jun–Aug. |
Flowering Jun–Jul. |
| Moist tundra and ridges, sand and gravel flats or beaches |
Clay flats, gravel, beaches, limestone outcrops, talus |
| 0-1000 m (0-3300 ft) |
0-500 m (0-1600 ft) |
|
LB; MB; NU; ON; QC; Greenland; Europe (Finland, Norway, Russia, Sweden)
|
NU; Greenland; Europe (Norway [Svalbard]) |
The synonymy above includes two North American names overlooked by C. L. Hitchcock (1941) and R. C. Rollins (1993). Draba alpina was broadly delimited by O. E. Schulz (1927) and included 17 varieties, some of which (e.g., corymbosa, oxycarpa, pilosa) are recognized herein as distinct species. The name D. alpina was so misapplied that it was used for any circumpolar or alpine, scapose, yellow-flowered, perennial Draba. Various chromosome numbers (e.g., 2n = 64, 80, 112, 120; S. I. Warwick and I. A. Al-Shehbaz 2006) have been reported for the species. As circumscribed here, it has the narrow distribution outlined above and includes plants with 2n = 80. Reports of the species from Alaska, Canadian Northwest Territories and Yukon, Siberia, eastern Asia, Russian Far East, and the Central Asian republics are either suspect or very unlikely. The entire D. alpina complex (including the above three species, D. glacialis Adams, D. macounii, etc.) is in need of critical molecular, cytological, and morphological study. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
As circumscribed here, Draba arctica consists of two distinct elements often recognized as subspecies (arctica and ostenfeldii; T. W. Böcher 1966); perhaps they are better treated as distinct species (D. arctica and D. ovibovina). Lack of adequate material and inability to examine all of the types in this complex prevent us from doing so. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
| FNA vol. 7, p. 290. |
FNA vol. 7, p. 291. |
| Brassicaceae > tribe Arabideae > Draba |
Brassicaceae > tribe Arabideae > Draba |
D. abajoensis, D. albertina, D. aleutica, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. cyclomorpha, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. helleriana, D. hitchcockii, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis |
D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. cyclomorpha, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. helleriana, D. hitchcockii, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis |
| D. alpina var. hydeana, D. alpina var. inflatisiliqua |
D. arctica subsp. ostenfeldii, D. arctica var. ostenfeldii, D. cinerea var. arctica, D. ostenfeldii, D. ostenfeldii var. ovibovina, D. ovibovina |
| Linnaeus: Sp. Pl. 2: 642. (1753) |
J. Vahl: in G. C. Oeder et al., Fl. Dan. 13(39): plate 5, 2294. (1840) |
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