Draba alpina |
Draba abajoensis |
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alpine Draba |
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Habit | Perennials; (cespitose); caudex branched (covered with persistent leaves or leaf remains); scapose. | Perennials; caudex simple or branched, (sometimes with persistent leaf bases); not scapose. |
Stems | unbranched, (0.3–)0.5–1.7(–2.8) dm, pubescent throughout, trichomes simple and 2-rayed, 0.3–0.8 mm, with 3–5-rayed ones, 0.1–0.3 mm. |
unbranched, (0.6–)1–2.5(–3.5) dm, pubescent proximally, usually glabrous distally, rarely sparsely pubescent, trichomes simple, 0.1–0.5 mm, often with stalked, 2(–4)-rayed ones. |
Basal leaves | rosulate; petiolate; petiole base (not thickened), ciliate, (trichomes simple, 0.3–1 mm); blade oblanceolate to obovate or lanceolate to oblong, 0.8–3(–4.5) cm × 2.5–6(–9) mm, margins entire, surfaces abaxially pubescent with stalked, 2–4-rayed trichomes, 0.1–0.5 mm, with simple ones (midvein obscure, not thickened), adaxially glabrous or sparsely pubescent with simple and stalked, 2–4-rayed trichomes. |
rosulate; petiolate; petiole (0.2–1(–2) cm), proximal margin ciliate, (trichomes simple, 0.2–0.7 mm); blade oblanceolate to spatulate, (0.6–)1–3.5(–4.3) cm × (2–)4–10(–14) mm, margins entire or dentate, surfaces pubescent with short-stalked, (2–)4-rayed trichomes, 0.05–0.35 mm, sometimes adaxially with simple ones. |
Cauline leaves | 0. |
(3–)5–9(–11); sessile; blade broadly ovate to lanceolate or oblong, margins entire or dentate, surfaces pubescent as basal or adaxially also with mostly simple trichomes. |
Racemes | 6–18-flowered, ebracteate, considerably elongated in fruit; rachis not flexuous, pubescent as stem. |
(11–)15–32(–47)-flowered, ebracteate, elongated in fruit; rachis not flexuous, glabrous or pubescent, trichomes simple and short-stalked, 2–4-rayed. |
Flowers | sepals (purplish tinged), narrowly ovate, 2.5–3 mm, pubescent, (trichomes simple and fewer, stalked, 2-rayed); petals bright yellow, narrowly obovate, 3.5–5 × 1.7–2.5 mm; anthers ovate, 0.3–0.4 mm. |
sepals ovate-oblong, 2.2–3.5 mm, subapically sparsely pubescent, (trichomes simple, short-stalked, and 2-rayed); petals yellow, oblanceolate, 4–6 × 1.5–2 mm; anthers oblong, 0.6–1 mm. |
Fruiting pedicels | ascending to divaricate-ascending, straight or, sometimes, slightly curved upwards, 4–14(–30) mm, pubescent, trichomes simple and 2–4-rayed. |
horizontal to divaricate-ascending, straight or curved, (5–)7–18(–23) mm, glabrous or sparsely pubescent abaxially, trichomes simple and 2-rayed. |
Fruits | elliptic, plane, flattened, 6–10 × 2–3 mm; valves glabrous or glabrescent, trichomes simple, (not confined to replum); ovules 12–24 per ovary; style 0.2–0.3 mm (stigma about as wide as style). |
elliptic to elliptic-lanceolate, plane, flattened, 5–9(–12) × 2–3 mm; valves glabrous; ovules 10–18 per ovary; style (0.8–)1.4–2.7(–3.6) mm. |
Seeds | (pale brown), ovoid, 0.9–1.3 × 0.6–0.9 mm. |
ovoid, 1–1.2 × 0.7–0.8 mm. |
2n | = 80. |
= 20. |
Draba alpina |
Draba abajoensis |
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Phenology | Flowering Jun–Aug. | Flowering May–Aug. |
Habitat | Moist tundra and ridges, sand and gravel flats or beaches | Spruce, fir, or pine forests, subalpine meadows |
Elevation | 0-1000 m (0-3300 ft) | 1900-3800 m (6200-12500 ft) |
Distribution |
LB; MB; NU; ON; QC; Greenland; Europe (Finland, Norway, Russia, Sweden)
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AZ; NM; UT |
Discussion | The synonymy above includes two North American names overlooked by C. L. Hitchcock (1941) and R. C. Rollins (1993). Draba alpina was broadly delimited by O. E. Schulz (1927) and included 17 varieties, some of which (e.g., corymbosa, oxycarpa, pilosa) are recognized herein as distinct species. The name D. alpina was so misapplied that it was used for any circumpolar or alpine, scapose, yellow-flowered, perennial Draba. Various chromosome numbers (e.g., 2n = 64, 80, 112, 120; S. I. Warwick and I. A. Al-Shehbaz 2006) have been reported for the species. As circumscribed here, it has the narrow distribution outlined above and includes plants with 2n = 80. Reports of the species from Alaska, Canadian Northwest Territories and Yukon, Siberia, eastern Asia, Russian Far East, and the Central Asian republics are either suspect or very unlikely. The entire D. alpina complex (including the above three species, D. glacialis Adams, D. macounii, etc.) is in need of critical molecular, cytological, and morphological study. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Plants of Draba abajoensis were treated by C. L. Hitchcock (1941), R. C. Rollins (1993), and N. H. Holmgren (2005b) as D. spectabilis. The differences between these taxa in chromosome number and trichome morphology strongly support their recognition as separate species (I. A. Al-Shehbaz and M. D. Windham 2007). Draba abajoensis has been collected from the Chuska and Lukachukai mountains in Apache County, Arizona, Chuska Mountains in San Juan County, New Mexico, and Abajo and La Sal mountains in San Juan and Grand counties, Utah. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 290. | FNA vol. 7, p. 288. |
Parent taxa | Brassicaceae > tribe Arabideae > Draba | Brassicaceae > tribe Arabideae > Draba |
Sibling taxa | ||
Synonyms | D. alpina var. hydeana, D. alpina var. inflatisiliqua | D. spectabilis var. glabrescens |
Name authority | Linnaeus: Sp. Pl. 2: 642. (1753) | Windham & Al-Shehbaz: Harvard Pap. Bot. 12: 416. (2007) |
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