Draba albertina |
Draba ruaxes |
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Alaska Draba, Alaska whitlow-grass, Alberta whitlow grass, slender Draba, slender whitlow-grass |
coast mountain Draba, coast mountain whitlow-grass, coast mountain whitlow-grass Draba, Rainier Draba |
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Habit | Annuals, biennials, or perennials; caudex (or base) simple or branched (poorly developed); rarely scapose. | Perennials; (cespitose, forming tufts); caudex branched (with persistent leaf bases, branches some terminating in sterile rosettes); scapose. |
Stems | branched distally, (0.3–)0.5–3(–4.2) dm, pubescent proximally, often glabrous distally, trichomes simple, 0.1–1 mm, sometimes with fewer, stalked, 2-rayed ones. |
unbranched, 0.2–0.6(–0.8) dm, often pubescent throughout, sometimes glabrate distally, trichomes simple, 0.4–1 mm, and 2–4-rayed, 0.1–0.4 mm. |
Basal leaves | rosulate; shortly petiolate; petiole (0–0.5 cm), ciliate throughout; blade obovate to oblanceolate or linear-lanceolate, (0.3–)1–2.8(–3.5) cm × (1–)2–6(–9) mm, margins entire or denticulate, (ciliate, trichomes simple, 0.4–1 mm), surfaces usually pubescent, abaxially with stalked, 2–4-rayed trichomes, 0.05–0.4(–0.5) mm, (rarely with simple trichomes along midvein), adaxially with simple trichomes sometimes also with 2-rayed ones, 0.07–0.4 mm, rarely glabrous. |
rosulate; subsessile; petiole base and margin ciliate, (trichomes simple, 0.4–1.4 mm); blade oblanceolate to obovate or suborbicular, 0.3–1 cm × 2–4.5 mm, margins entire, surfaces pubescent, abaxially with stalked, 2–10-rayed, stellate trichomes, 0.2–0.8 mm, adaxially with simple trichomes, 0.4–1 mm, sometimes with smaller, 2–4-rayed ones, (midvein obscure). |
Cauline leaves | (0 or) 1–3(–5); sessile; blade lanceolate to elliptic or ovate, margins entire or denticulate, surfaces pubescent as basal. |
0. |
Racemes | (2–)6–30(–50)-flowered, ebracteate, elongated in fruit; rachis straight or, rarely, flexuous, glabrous. |
(2–)4–10(–14)-flowered, ebracteate, elongated in fruit; rachis not flexuous, pubescent as stem or glabrous. |
Flowers | (chasmogamous, petaliferous); sepals ovate, 1.4–2.1 mm, glabrous or sparsely pubescent, (trichomes simple); petals yellow, spatulate to oblanceolate, 2–3.2 × 0.7–1.2 mm; anthers ovate, 0.15–0.25 mm. |
sepals ovate, 2–3 mm, pubescent, (trichomes simple with fewer, 2-rayed ones); petals bright yellow, obovate, 4–6 × 2–3.5 mm; anthers oblong, 0.4–0.5 mm. |
Fruiting pedicels | divaricate-ascending or horizontal, (not appressed to rachis), usually straight, rarely curved upward, (3–)5–14(–16) mm (subequaling or shorter than fruit), usually glabrous, rarely sparsely puberulent. |
ascending to divaricate-ascending, straight or slightly curved upward, 3–7(–9) mm, pubescent or glabrous, trichomes simple and 2–4-rayed. |
Fruits | lanceolate to narrowly elliptic or linear, plane, flattened, (4–)6–12(–15) × (1–)1.4–2.1 mm; valves glabrous; ovules (20–)24–38(–44) per ovary; style 0.01–0.12 mm. |
elliptic to lanceolate or ovate to suborbicular, plane, flattened, 4–8(–10) × 3–4.5 mm; valves puberulent, trichomes simple with fewer 2-rayed ones, 0.1–0.35 mm; ovules 12–16 per ovary; style 0.5–0.9(–1.1) mm. |
Seeds | oblong, 0.7–1 × 0.4–0.5 mm. |
oblong, 1.5–2 × 1–1.3 mm. |
2n | = 24. |
= 72. |
Draba albertina |
Draba ruaxes |
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Phenology | Flowering Jun–Aug. | Flowering Jun–Jul. |
Habitat | Open woodlands, pine forests, meadows, rocky knolls, alpine slopes, stream banks, disturbed areas | Rock outcrops, talus slopes, ridges, alpine summits |
Elevation | 900-3700 m (3000-12100 ft) | 500-2400 m (1600-7900 ft) |
Distribution |
AK; AZ; CA; CO; ID; MT; NM; NV; OR; UT; WA; WY; AB; BC; NT; YT
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AK; WA; AB; BC; YT
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Discussion | O. E. Schulz (1927) and C. L. Hitchcock (1941) confused the limits of Draba albertina, D. crassifolia, and D. stenoloba; the latter author treated the first two species as conspecific. G. A. Mulligan (1975) demonstrated that the three taxa are chromosomally and morphologically distinct (see D. crassifolia and D. stenoloba for differences from D. albertina). The ranges of D. albertina and D. crassifolia overlap extensively, and occasional sterile hybrids are encountered. The Alaskan record of Draba albertina is based on Minard 4 (ALA), collected on the northern coast of Afognak Island (58°22’N, 152°28’W). The record from New Mexico is based on O’Kane & Hedin 3871 (ISTC, SJC), collected in Chuska Mountains (36°8’11”N, 108°54’19”W). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
C. L. Hitchcock (1941) treated Draba ruaxes as a variety of D. ventosa; as demonstrated by G. A. Mulligan (1971b), the two are quite distinct. Draba ruaxes is an outcrossing hexaploid with well-formed anthers and pollen, and abundant, simple trichomes on leaves, stems, sepals, and fruits. By contrast, D. ventosa is an apomictic triploid with abortive anthers and/or pollen, and no simple trichomes anywhere on the plant. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 288. | FNA vol. 7, p. 334. |
Parent taxa | Brassicaceae > tribe Arabideae > Draba | Brassicaceae > tribe Arabideae > Draba |
Sibling taxa | ||
Synonyms | D. crassifolia var. albertina, D. crassifolia var. nevadensis, D. deflexa, D. nitida, D. nitida var. nana, D. stenoloba var. nana, D. stenoloba var. ramosa | D. ventosa var. ruaxes |
Name authority | Greene: Pittonia 4: 312. (1901) | Payson & H. St. John: Proc. Biol. Soc. Wash. 43: 117. (1930) |
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