Draba albertina |
Draba grandis |
|
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Alaska Draba, Alaska whitlow-grass, Alberta whitlow grass, slender Draba, slender whitlow-grass |
north Pacific Draba |
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Habit | Annuals, biennials, or perennials; caudex (or base) simple or branched (poorly developed); rarely scapose. | Perennials; caudex simple or branched (fleshy, with persistent leaf bases); not scapose. |
Stems | branched distally, (0.3–)0.5–3(–4.2) dm, pubescent proximally, often glabrous distally, trichomes simple, 0.1–1 mm, sometimes with fewer, stalked, 2-rayed ones. |
(decumbent), unbranched, (0.2–)0.5–2.7(–3.7) dm, pubescent throughout, trichomes 2–4-rayed, 0.05–0.2 mm. |
Basal leaves | rosulate; shortly petiolate; petiole (0–0.5 cm), ciliate throughout; blade obovate to oblanceolate or linear-lanceolate, (0.3–)1–2.8(–3.5) cm × (1–)2–6(–9) mm, margins entire or denticulate, (ciliate, trichomes simple, 0.4–1 mm), surfaces usually pubescent, abaxially with stalked, 2–4-rayed trichomes, 0.05–0.4(–0.5) mm, (rarely with simple trichomes along midvein), adaxially with simple trichomes sometimes also with 2-rayed ones, 0.07–0.4 mm, rarely glabrous. |
rosulate; long-petiolate; petiole (winged, (1–)4–15 cm), often not ciliate (or ciliate to blade apex, trichomes simple, 0.3–0.8 mm); blade (somewhat fleshy), oblanceolate to spatulate or obovate, (1–)2–11(–17) cm × (5–)8–30(–45) mm, margins often coarsely dentate, (pubescent as petiole), surfaces usually pubescent, abaxially with stalked, 2–4-rayed trichomes, 0.05–0.25 mm, adaxially similar, or also with simple and long-stalked, 2-rayed trichomes, to 0.8 mm, rarely glabrescent, with mostly simple and 2-rayed trichomes. |
Cauline leaves | (0 or) 1–3(–5); sessile; blade lanceolate to elliptic or ovate, margins entire or denticulate, surfaces pubescent as basal. |
2–12(–16); sessile or petiolate; blade oblanceolate to obovate, margins dentate or entire, surfaces pubescent as basal. |
Racemes | (2–)6–30(–50)-flowered, ebracteate, elongated in fruit; rachis straight or, rarely, flexuous, glabrous. |
5–26(–32)-flowered, ebracteate, elongated in fruit; rachis not flexuous, pubescent as stem. |
Flowers | (chasmogamous, petaliferous); sepals ovate, 1.4–2.1 mm, glabrous or sparsely pubescent, (trichomes simple); petals yellow, spatulate to oblanceolate, 2–3.2 × 0.7–1.2 mm; anthers ovate, 0.15–0.25 mm. |
sepals broadly ovate, 3–4 mm, pubescent, (trichomes simple and short-stalked, 2-rayed); petals yellow, oblanceolate to spatulate, 4.5–7 × 1.8–3 mm; anthers oblong, 0.7–1 mm. |
Fruiting pedicels | divaricate-ascending or horizontal, (not appressed to rachis), usually straight, rarely curved upward, (3–)5–14(–16) mm (subequaling or shorter than fruit), usually glabrous, rarely sparsely puberulent. |
horizontal to divaricate-ascending or ascending, usually straight, rarely curved upward, (5–)10–22(–27) mm, pubescent, trichomes 2–4-rayed (0.05–0.3 mm), sometimes with simple and spurred ones. |
Fruits | lanceolate to narrowly elliptic or linear, plane, flattened, (4–)6–12(–15) × (1–)1.4–2.1 mm; valves glabrous; ovules (20–)24–38(–44) per ovary; style 0.01–0.12 mm. |
oblong to lanceolate, or ovate to suborbicular, slightly twisted or plane, flattened, (6–)10–20(–25) × 4–7(–9) mm; valves glabrous; ovules 24–52 per ovary; style (0.2–)0.4–1.6(–2) mm. |
Seeds | oblong, 0.7–1 × 0.4–0.5 mm. |
ovoid, 1.4–2 × 0.8–1.2 mm. |
2n | = 24. |
= 36. |
Draba albertina |
Draba grandis |
|
Phenology | Flowering Jun–Aug. | Flowering May–Jul. |
Habitat | Open woodlands, pine forests, meadows, rocky knolls, alpine slopes, stream banks, disturbed areas | Rocky bluffs above salt-water beaches, loamy seaside banks, sea bird rookeries, coastal herbaceous tundra and sandy blowouts |
Elevation | 900-3700 m (3000-12100 ft) | 0-50(-200) m (0-200(-700) ft) |
Distribution |
AK; AZ; CA; CO; ID; MT; NM; NV; OR; UT; WA; WY; AB; BC; NT; YT
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AK; BC; e Asia (Kuril and Ratmanov islands, Russian Far East) |
Discussion | O. E. Schulz (1927) and C. L. Hitchcock (1941) confused the limits of Draba albertina, D. crassifolia, and D. stenoloba; the latter author treated the first two species as conspecific. G. A. Mulligan (1975) demonstrated that the three taxa are chromosomally and morphologically distinct (see D. crassifolia and D. stenoloba for differences from D. albertina). The ranges of D. albertina and D. crassifolia overlap extensively, and occasional sterile hybrids are encountered. The Alaskan record of Draba albertina is based on Minard 4 (ALA), collected on the northern coast of Afognak Island (58°22’N, 152°28’W). The record from New Mexico is based on O’Kane & Hedin 3871 (ISTC, SJC), collected in Chuska Mountains (36°8’11”N, 108°54’19”W). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Almost all recent North American authors have used the name Draba hyperborea for this species. A. N. Berkutenko (1995) clearly showed that the type of that name belongs to an entirely different species that she placed in the genus Schivereckia Andrzejowski ex de Candolle. Draba grandis thus becomes the correct name for the North American taxon. Except for its fleshy leaves, Nesodraba is indistinguishable morphologically and molecularly from other species of Draba. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 288. | FNA vol. 7, p. 309. |
Parent taxa | Brassicaceae > tribe Arabideae > Draba | Brassicaceae > tribe Arabideae > Draba |
Sibling taxa | ||
Synonyms | D. crassifolia var. albertina, D. crassifolia var. nevadensis, D. deflexa, D. nitida, D. nitida var. nana, D. stenoloba var. nana, D. stenoloba var. ramosa | Cochlearia siliquosa, Cochlearia spathulata, D. greenei, D. hatchiae, D. hyperborea var. spathulata, D. spathulata, Nesodraba grandis, Nesodraba megalocarpa, Nesodraba siliquosa |
Name authority | Greene: Pittonia 4: 312. (1901) | Langsdorff ex de Candolle: Syst. Nat. 2: 355. (1821) |
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