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cliff dwarf-primrose, smooth douglasia

Bering dwarf-primrose

Habit Plants loosely cespitose mats with branched caudex. Plants loosely cespitose cushions or mats, each with branched caudex.
Stems

prostrate, loosely covered with marcescent, gray to light brown leaves (becoming remote in age).

prostrate to ascending, with marcescent, imbricate, reddish leaves proximally and terminal, green leaf rosettes.

Leaves

spreading, thin;

blade oblong-lanceolate or oblanceolate to spatulate, 5–20 × 2–6 mm, margins entire or slightly toothed, sometimes ciliolate, hairs simple, apex obtuse to slightly acute, surfaces glabrous.

prominently ascending, thin;

blade linear to oblong, 3–10 × 1–1.5 mm, margins entire or toothed, apex obtuse to slightly acute, surfaces densely covered with minute, branched hairs.

Scapes

2–7 mm, elongating little in fruit, minutely hairy, hairs stellate and branched.

1–10 mm in early anthesis, to ca. 6 cm in fruit, densely hairy, hairs branched and stellate.

Inflorescences

2–10-flowered, bracteate;

bracts 3–8, lanceolate to ovate, 3–8 × 1–3 mm, glabrous or sparsely hairy, hairs minute, branched.

1(–2)-flowered, usually bracteate, sometimes ebracteate;

bract 1, lanceolate to lanceolate-ovate, 2–3 × 0.5–1 mm, hairy, hairs stellate.

Pedicels

2–15 mm.

absent.

Flowers

calyx 6–7 × 3–4 mm, stellate-pubescent;

corolla rose-pink, violet in age, limb 8–15 mm diam., lobes 3–4 × 2–3 mm, margins entire or erose.

calyx 3–5 × 3–4 mm, densely hairy, hairs minute, stellate and branched;

corolla pink, white in age, limb 5–7 mm diam., lobes 2 × 1 mm, margins entire or slightly erose.

Douglasia laevigata

Douglasia beringensis

Phenology Flowering early summer. Flowering early summer.
Habitat Rocky areas Gravel slopes and outcrops
Elevation 30-2000 m (100-6600 ft) 300-500 m (1000-1600 ft)
Distribution
from FNA
OR; WA; BC
[WildflowerSearch map]
[BONAP county map]
from FNA
AK
[BONAP county map]
Discussion

Although the first collection of Douglasia laevigata was from the “Mountains near Mt. Hood,” the original description of the species was based on plants collected in the Columbia River gorge, which thus represent the nomenclaturally typical variety (L. Constance 1938), even though that entity constitutes an ecological variant with almost glabrous leaves and loose umbels known only from the gorge. The widespread form, var. ciliolata, has more compact umbels and larger, more toothed, conspicuously ciliolate leaves. Because intermediate forms occur commonly, and even the type specimen of D. laevigata has cilia, the infraspecific taxa are not recognized here.

A population of Douglasia laevigata from Cone Peak in the Cascade Mountains of Oregon has been reported as heterostylous due to the occurrence of a protruding “pin type” stigma from the corolla throat; this represents only a local stigmatic anomaly functioning as minor spatial separation of anthers and stigma in early anthesis rather than true heterostyly as seen in Primula. No stigmatic or pollen dimorphisms occur in these plants and no parallel “thrum type” flowers are known.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Douglasia beringensis has been found in western Alaska from the Seward Peninsula to the western Alaska Range; a fragmentary collection from the Yukon River in central Alaska is questionable. It differs from other northern species of Douglasia in the pervasive stellate hairs covering all vegetative parts. Douglasia gormanii has sparser forked hairs only on leaves and peduncles, not calyces; plants of D. gormanii also lack the silvery aspect that is characteristic of D. beringensis from its stellate indument.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 8, p. 266. FNA vol. 8, p. 265.
Parent taxa Primulaceae > Douglasia Primulaceae > Douglasia
Sibling taxa
D. alaskana, D. arctica, D. beringensis, D. gormanii, D. idahoensis, D. montana, D. nivalis, D. ochotensis
D. alaskana, D. arctica, D. gormanii, D. idahoensis, D. laevigata, D. montana, D. nivalis, D. ochotensis
Synonyms D. laevigata subsp. ciliolata, D. laevigata var. ciliolata
Name authority A. Gray: Proc. Amer. Acad. Arts 16: 105. 1880 , S. Kelso: Novon 4: 381, fig. 1. (1994)
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