FNA: Diplacus viscidus vs. Diplacus calycinus

	Diplacus viscidus	Diplacus calycinus
	sticky monkeyflower, viscid monkeyflower	Kaweah River bush monkeyflower, rock bush monkeyflower
Habit	Herbs, annual.	Subshrubs.
Stems	erect, (30–)60–370 mm, densely glandular-pubescent with viscid hairs.	erect, 150–1500 mm, glandular-pubescent to viscid-villous.
Leaves	usually cauline, relatively even-sized or largest proximally and gradually reduced distally; petiole absent; blade obovate to narrowly elliptic, (4–)8–54(–70) × (2–)3–23 mm, margins entire or serrate, plane, apex obtuse to rounded, surfaces: proximals glabrous abaxially, distals glandular-pubescent.	cauline, relatively even-sized; petiole absent; blade elliptic-lanceolate or lanceolate to narrowly lanceolate or linear-lanceolate, sometimes narrowly oblong, 20–75(–100) × 4–20(–28) mm, margins entire or shallowly crenate, plane or revolute, apex acute to obtuse, abaxial surfaces moderately villous, hairs unbranched, vitreous, adaxial glabrous.
Pedicels	1–4(–5) mm in fruit.	3–5 mm in fruit.
Flowers	1 per node, chasmogamous.	2 per node, chasmogamous.
Styles	glandular-puberulent.	minutely glandular.
Corollas	lavender to magenta with diffuse dark markings on sides of darker tube-throat and with dark red-purple midveins on lobes extending from throat, lobes not dark at base, floor white or yellow, fading to white at mouth, palate ridges white or yellow fading to white distally, throat ceiling glabrous, tube-throat 10–20 mm, limb 8–20 mm diam., not bilabiate.	usually pale yellow or cream to yellow, not spotted or striped, palate ridges yellow to golden yellow, tube-throat 35–42 mm, limb 20–30 mm diam., bilabiate, lobes oblong, apex of adaxial 2 each shallowly, asymmetrically incised.
Calyces	symmetrically attached to pedicels, inflated in fruit, (7–)8–15 mm, villous, hairs eglandular, lobes subequal, apex acute to attenuate, ribs and intercostal areas often reddish.	not inflated in fruit, 28–40 mm, densely glandular-pubescent to short glandular-villous, tube slightly dilated distally, lobes unequal, apex acute, ribs green, intercostal areas light green.
Capsules	7–11 mm.	25–35 mm.
Anthers	included, ciliate.	exserted, glabrous.
Stigmas	included, lobes unequal, abaxial 1.5 times adaxial.	exserted, lobes equal.
2n	= 16.	= 20.

	Diplacus viscidus	Diplacus calycinus
Phenology	Flowering Apr–Jul.	Flowering Apr–Jul.
Habitat	Chaparral clearings and openings.	Granite outcrops, boulders, rocky gullies.
Elevation	90–1300 m. (300–4300 ft.)	(300–)700–2200 m. ((1000–)2300–7200 ft.)
Distribution	CA [WildflowerSearch map] [BONAP county map]	CA [WildflowerSearch map] [BONAP county map]
Discussion	D. M. Thompson (2005) noted that Diplacus viscidus and D. compactus (as Mimulus viscidus var. compactus) are parapatric and may intergrade in central Mariposa County. The two taxa are distinguished by the presence or absence of dark stripes on the corolla lobe midveins, which are evident even on herbarium specimens. Thompson found that the two remained distinct when grown together in the greenhouse. Diplacus viscidus is known from Amador, Calaveras, Eldorado, Mariposa, Merced, and Tuolumne counties; D. compactus continues south through Fresno, northern Kern, Madera, Mariposa, and Tulare counties. (Discussion copyrighted by Flora of North America; reprinted with permission.)	Although first described as a separate species, Diplacus calycinus has more recently been treated at subspecific or varietal rank (A. L. Grant 1924; F. W. Pennell 1951; P. A. Munz and D. D. Keck 1973). D. M. Thompson (2005) included both D. calycinus and D. longiflorus within his concept of Mimulus aurantiacus var. pubescens (Torrey) D. M. Thompson. He did not reference the study of sect. Diplacus by M. C. Tulig (2000), but results from the Tulig morphometric analyses indicated that D. calycinus is distinct from D. longiflorus, especially in corolla length, corolla tube length, and style length. The type of Diplacus calycinus is from Tulare County, and the concept of the species is perhaps best restricted to the Sierran population system in Fresno, Kern, and Tulare counties, disjunct from D. longiflorus, which occurs primarily in coastal counties. The Sierran system is characterized by distinct abaxial leaf vestiture; the hairs are unbranched, broad, and vitreous, compared to the branched, thinner, and dull hairs of D. longiflorus. Plants of D. calycinus parapatric with D. longiflorus also show a tendency toward the characteristic vestiture and also have lighter-colored (but more variable in color) corollas with narrower but slightly shorter tubes. Intergradation between D. calycinus and D. longiflorus occurs in the region connecting the San Bernardino and San Gabriel mountains in San Bernardino County. (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 17, p. 432.	FNA vol. 17, p. 449.
Parent taxa	Phrymaceae > Diplacus	Phrymaceae > Diplacus
Sibling taxa	D. angustatus, D. aridus, D. aurantiacus, D. bicolor, D. bigelovii, D. bolanderi, D. brandegeei, D. brevipes, D. calycinus, D. cascadensis, D. clevelandii, D. clivicola, D. compactus, D. congdonii, D. constrictus, D. cusickii, D. cusickioides, D. deschutesensis, D. douglasii, D. fremontii, D. grandiflorus, D. graniticola, D. jepsonii, D. johnstonii, D. kelloggii, D. layneae, D. leptaleus, D. linearis, D. longiflorus, D. mephiticus, D. mohavensis, D. nanus, D. ovatus, D. parryi, D. parviflorus, D. pictus, D. pulchellus, D. puniceus, D. pygmaeus, D. rattanii, D. rupicola, D. rutilus, D. thompsonii, D. torreyi, D. traskiae, D. tricolor, D. vandenbergensis	D. angustatus, D. aridus, D. aurantiacus, D. bicolor, D. bigelovii, D. bolanderi, D. brandegeei, D. brevipes, D. cascadensis, D. clevelandii, D. clivicola, D. compactus, D. congdonii, D. constrictus, D. cusickii, D. cusickioides, D. deschutesensis, D. douglasii, D. fremontii, D. grandiflorus, D. graniticola, D. jepsonii, D. johnstonii, D. kelloggii, D. layneae, D. leptaleus, D. linearis, D. longiflorus, D. mephiticus, D. mohavensis, D. nanus, D. ovatus, D. parryi, D. parviflorus, D. pictus, D. pulchellus, D. puniceus, D. pygmaeus, D. rattanii, D. rupicola, D. rutilus, D. thompsonii, D. torreyi, D. traskiae, D. tricolor, D. vandenbergensis, D. viscidus
Synonyms	Mimulus viscidus, M. fremontii var. viscidus, M. subsecundus var. viscidus	D. longiflorus var. calycinus, Mimulus longiflorus subsp. calycinus, M. longiflorus var. calycinus
Name authority	(Congdon) G. L. Nesom: Phytoneuron 2012-39: 29. (2012)	Eastwood: Bot. Gaz. 41: 287. (1906)
Web links	Local floras: CA Local Web sites: CalFlora, CalPhotos WildflowerSearch iNaturalist (observations) LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	Local floras: CA Local Web sites: CalFlora, CalPhotos WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Diplacus calycinus

sticky monkeyflower, viscid monkeyflower

Kaweah River bush monkeyflower, rock bush monkeyflower

Habit

Herbs, annual.

Subshrubs.

Stems

erect, (30–)60–370 mm, densely glandular-pubescent with viscid hairs.

erect, 150–1500 mm, glandular-pubescent to viscid-villous.

Leaves

usually cauline, relatively even-sized or largest proximally and gradually reduced distally;

petiole absent;

blade obovate to narrowly elliptic, (4–)8–54(–70) × (2–)3–23 mm, margins entire or serrate, plane, apex obtuse to rounded, surfaces: proximals glabrous abaxially, distals glandular-pubescent.

cauline, relatively even-sized;

petiole absent;

blade elliptic-lanceolate or lanceolate to narrowly lanceolate or linear-lanceolate, sometimes narrowly oblong, 20–75(–100) × 4–20(–28) mm, margins entire or shallowly crenate, plane or revolute, apex acute to obtuse, abaxial surfaces moderately villous, hairs unbranched, vitreous, adaxial glabrous.

Pedicels

1–4(–5) mm in fruit.

3–5 mm in fruit.

Flowers

1 per node, chasmogamous.

2 per node, chasmogamous.

Styles

glandular-puberulent.

minutely glandular.

Corollas

lavender to magenta with diffuse dark markings on sides of darker tube-throat and with dark red-purple midveins on lobes extending from throat, lobes not dark at base, floor white or yellow, fading to white at mouth, palate ridges white or yellow fading to white distally, throat ceiling glabrous, tube-throat 10–20 mm, limb 8–20 mm diam., not bilabiate.

usually pale yellow or cream to yellow, not spotted or striped, palate ridges yellow to golden yellow, tube-throat 35–42 mm, limb 20–30 mm diam., bilabiate, lobes oblong, apex of adaxial 2 each shallowly, asymmetrically incised.

Calyces

symmetrically attached to pedicels, inflated in fruit, (7–)8–15 mm, villous, hairs eglandular, lobes subequal, apex acute to attenuate, ribs and intercostal areas often reddish.

not inflated in fruit, 28–40 mm, densely glandular-pubescent to short glandular-villous, tube slightly dilated distally, lobes unequal, apex acute, ribs green, intercostal areas light green.

Capsules

7–11 mm.

25–35 mm.

Anthers

included, ciliate.

exserted, glabrous.

Stigmas

included, lobes unequal, abaxial 1.5 times adaxial.

exserted, lobes equal.

= 16.

= 20.

Diplacus viscidus

Diplacus calycinus

Phenology

Flowering Apr–Jul.

Habitat

Chaparral clearings and openings.

Granite outcrops, boulders, rocky gullies.

Elevation

90–1300 m. (300–4300 ft.)

(300–)700–2200 m. ((1000–)2300–7200 ft.)

Distribution

[WildflowerSearch map]

[BONAP county map]

[WildflowerSearch map]

[BONAP county map]

Discussion

D. M. Thompson (2005) noted that Diplacus viscidus and D. compactus (as Mimulus viscidus var. compactus) are parapatric and may intergrade in central Mariposa County. The two taxa are distinguished by the presence or absence of dark stripes on the corolla lobe midveins, which are evident even on herbarium specimens. Thompson found that the two remained distinct when grown together in the greenhouse.

Diplacus viscidus is known from Amador, Calaveras, Eldorado, Mariposa, Merced, and Tuolumne counties; D. compactus continues south through Fresno, northern Kern, Madera, Mariposa, and Tulare counties.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Although first described as a separate species, Diplacus calycinus has more recently been treated at subspecific or varietal rank (A. L. Grant 1924; F. W. Pennell 1951; P. A. Munz and D. D. Keck 1973). D. M. Thompson (2005) included both D. calycinus and D. longiflorus within his concept of Mimulus aurantiacus var. pubescens (Torrey) D. M. Thompson. He did not reference the study of sect. Diplacus by M. C. Tulig (2000), but results from the Tulig morphometric analyses indicated that D. calycinus is distinct from D. longiflorus, especially in corolla length, corolla tube length, and style length.

The type of Diplacus calycinus is from Tulare County, and the concept of the species is perhaps best restricted to the Sierran population system in Fresno, Kern, and Tulare counties, disjunct from D. longiflorus, which occurs primarily in coastal counties. The Sierran system is characterized by distinct abaxial leaf vestiture; the hairs are unbranched, broad, and vitreous, compared to the branched, thinner, and dull hairs of D. longiflorus. Plants of D. calycinus parapatric with D. longiflorus also show a tendency toward the characteristic vestiture and also have lighter-colored (but more variable in color) corollas with narrower but slightly shorter tubes. Intergradation between D. calycinus and D. longiflorus occurs in the region connecting the San Bernardino and San Gabriel mountains in San Bernardino County.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 17, p. 432.

FNA vol. 17, p. 449.

Parent taxa

Phrymaceae > Diplacus

Sibling taxa

D. angustatus, D. aridus, D. aurantiacus, D. bicolor, D. bigelovii, D. bolanderi, D. brandegeei, D. brevipes, D. calycinus, D. cascadensis, D. clevelandii, D. clivicola, D. compactus, D. congdonii, D. constrictus, D. cusickii, D. cusickioides, D. deschutesensis, D. douglasii, D. fremontii, D. grandiflorus, D. graniticola, D. jepsonii, D. johnstonii, D. kelloggii, D. layneae, D. leptaleus, D. linearis, D. longiflorus, D. mephiticus, D. mohavensis, D. nanus, D. ovatus, D. parryi, D. parviflorus, D. pictus, D. pulchellus, D. puniceus, D. pygmaeus, D. rattanii, D. rupicola, D. rutilus, D. thompsonii, D. torreyi, D. traskiae, D. tricolor, D. vandenbergensis

D. angustatus, D. aridus, D. aurantiacus, D. bicolor, D. bigelovii, D. bolanderi, D. brandegeei, D. brevipes, D. cascadensis, D. clevelandii, D. clivicola, D. compactus, D. congdonii, D. constrictus, D. cusickii, D. cusickioides, D. deschutesensis, D. douglasii, D. fremontii, D. grandiflorus, D. graniticola, D. jepsonii, D. johnstonii, D. kelloggii, D. layneae, D. leptaleus, D. linearis, D. longiflorus, D. mephiticus, D. mohavensis, D. nanus, D. ovatus, D. parryi, D. parviflorus, D. pictus, D. pulchellus, D. puniceus, D. pygmaeus, D. rattanii, D. rupicola, D. rutilus, D. thompsonii, D. torreyi, D. traskiae, D. tricolor, D. vandenbergensis, D. viscidus

Synonyms

Mimulus viscidus, M. fremontii var. viscidus, M. subsecundus var. viscidus

D. longiflorus var. calycinus, Mimulus longiflorus subsp. calycinus, M. longiflorus var. calycinus

Name authority

(Congdon) G. L. Nesom: Phytoneuron 2012-39: 29. (2012)

Eastwood: Bot. Gaz. 41: 287. (1906)

Web links

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

Choose a data source: Flora of North America

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Diplacus viscidus

Diplacus calycinus

Diplacus viscidus

Diplacus calycinus