FNA: Diplacus rattanii vs. Diplacus cusickii

	Diplacus rattanii	Diplacus cusickii
	Rattan's monkeyflower	Cusick's monkey-flower
Habit	Herbs, annual.	Herbs, annual.
Stems	erect, 10–180(–230) mm, densely glandular-pubescent and viscid.	erect to erect-ascending, 10–80 mm, distal internodes 2–20 mm, minutely glandular-puberulent.
Leaves	basal and cauline, basal in rosette, cauline gradually reduced distally; petiole absent; blade obovate to narrowly elliptic, 3–46(–70) × 1–20(–25) mm, margins entire or crenate, plane, apex rounded or obtuse, surfaces: proximals glabrate, distals glandular-pubescent and viscid.	basal and cauline or usually cauline, relatively even-sized or gradually larger distally; petiole absent, proximal base short petiole-like, 1–5 mm; blade ovate to broadly elliptic-ovate, (10–)15–25(–35) × 4–17 mm, margins entire, plane, apex acuminate, surfaces glabrous or adaxial minutely glandular-puberulent.
Pedicels	1–3(–6) mm in fruit.	1–1.5 mm in fruit.
Flowers	1 per node, chasmogamous.	2 per node, or 1 or 2 per node on 1 plant, chasmogamous.
Styles	eglandular-puberulent.	pubescent, at least on distal 1/2.
Corollas	pink to magenta, throat floor with 3 dark purple lines meeting abaxial lip lobes, palate ridges yellow, tube-throat 7–10 mm, limb 4–7 mm diam., not bilabiate.	magenta or rose purple, tube yellow, throat yellow, throat and distal tube red-spotted on floor, palate ridges yellow, tube-throat 13–16(–19) mm, limb 16–24 mm diam., bilabiate.
Calyces	symmetrically attached to pedicels, inflated in fruit, 5–10 mm, glandular-pubescent and viscid, lobes subequal, apex obtuse, often apiculate, ribs dark green to purplish, intercostal areas whitish.	symmetrically attached to pedicels, not inflated in fruit, 7–10 mm, glabrous or minutely stipitate-glandular, lobes unequal, apex linear-acuminate, sharp-pointed, ribs green distally, intercostal areas whitish.
Capsules	7–11(–12) mm.	10–15 mm.
Anthers	nearly exserted, glabrous.	included, glabrous or sparsely hirsutulous.
Stigmas	nearly exserted, lobes unequal, abaxial 5–7 times adaxial.	exserted, lobes subequal.
2n	= 16.

	Diplacus rattanii	Diplacus cusickii
Phenology	Flowering Apr–Jul.	Flowering May–Jul.
Habitat	Recently burned or cleared areas, sandhills, sandstone outcrops, sandy gravel and loam, decomposed granite, serpentine-derived soils, open chaparral, chaparral margins, open yellow pine-manzanita woodlands.	Slopes, canyons, washes, ditches, sand talus, diatomaceous slopes, basalt outcrops, black volcanic gravel, volcanic ash and sand, sagebrush areas.
Elevation	300–1300 m. (1000–4300 ft.)	800–1000 m. (2600–3300 ft.)
Distribution	CA [WildflowerSearch map]	ID; OR [WildflowerSearch map] [BONAP county map]
Discussion	Diplacus rattanii occurs mostly in near-coastal localities from Glenn and Lake counties south to Ventura County. (Discussion copyrighted by Flora of North America; reprinted with permission.)	Diplacus cusickii is endemic to northern Malheur County, Oregon, and along the Snake River in Ada and Owyhee counties, Idaho. Its narrow geographic range reflects the segregation of the more widely distributed D. cusickioides. D. M. Thompson (2005) noted that collections in northern Malheur County, Oregon, were intermediate between Mimulus cusickii and M. nanus; the type of M. cusickii is from this area and is among the narrowly endemic, supposedly putative intermediates (G. L. Nesom 2013c). These plants have abruptly and sharply acuminate leaf apices like the more widespread form traditionally identified as Diplacus cusickii; they differ in having glabrous leaf surfaces. Typical D. nanus occurs in close sympatry, without intergradation, with the populations in northern Malheur County. Because of their distinctive morphology and coherent geography, the northern Malheur County plants are reasonably recognized as a distinct species. The more widely distributed form formerly identified as D. cusickii now is identified as D. cusickioides. Populations of D. cusickii in the narrow sense along the Snake River in Ada and Owyhee counties, Idaho, may have dispersed there from the Oregon center. Some plants of D. cusickioides in the Leslie Gulch area of east-central Malheur County have somewhat reduced vestiture, approaching that of D. cusickii. (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 17, p. 432.	FNA vol. 17, p. 439.
Parent taxa	Phrymaceae > Diplacus	Phrymaceae > Diplacus
Sibling taxa	D. angustatus, D. aridus, D. aurantiacus, D. bicolor, D. bigelovii, D. bolanderi, D. brandegeei, D. brevipes, D. calycinus, D. cascadensis, D. clevelandii, D. clivicola, D. compactus, D. congdonii, D. constrictus, D. cusickii, D. cusickioides, D. deschutesensis, D. douglasii, D. fremontii, D. grandiflorus, D. graniticola, D. jepsonii, D. johnstonii, D. kelloggii, D. layneae, D. leptaleus, D. linearis, D. longiflorus, D. mephiticus, D. mohavensis, D. nanus, D. ovatus, D. parryi, D. parviflorus, D. pictus, D. pulchellus, D. puniceus, D. pygmaeus, D. rupicola, D. rutilus, D. thompsonii, D. torreyi, D. traskiae, D. tricolor, D. vandenbergensis, D. viscidus	D. angustatus, D. aridus, D. aurantiacus, D. bicolor, D. bigelovii, D. bolanderi, D. brandegeei, D. brevipes, D. calycinus, D. cascadensis, D. clevelandii, D. clivicola, D. compactus, D. congdonii, D. constrictus, D. cusickioides, D. deschutesensis, D. douglasii, D. fremontii, D. grandiflorus, D. graniticola, D. jepsonii, D. johnstonii, D. kelloggii, D. layneae, D. leptaleus, D. linearis, D. longiflorus, D. mephiticus, D. mohavensis, D. nanus, D. ovatus, D. parryi, D. parviflorus, D. pictus, D. pulchellus, D. puniceus, D. pygmaeus, D. rattanii, D. rupicola, D. rutilus, D. thompsonii, D. torreyi, D. traskiae, D. tricolor, D. vandenbergensis, D. viscidus
Synonyms	Mimulus rattanii, M. rattanii subsp. decurtatus	Eunanus cusickii, Mimulus cusickii
Name authority	(A. Gray) G. L. Nesom: Phytoneuron 2012-39: 29. (2012)	(Greene) G. L. Nesom: Phytoneuron 2012-39: 28. (2012)
Web links	Local floras: CA Local Web sites: CalFlora, CalPhotos WildflowerSearch iNaturalist (observations) LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	Local floras: OR Local Web sites: Flora NW, PNW Herbaria WildflowerSearch iNaturalist (observations) LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Diplacus cusickii

Rattan's monkeyflower

Cusick's monkey-flower

Habit

Herbs, annual.

Stems

erect, 10–180(–230) mm, densely glandular-pubescent and viscid.

erect to erect-ascending, 10–80 mm, distal internodes 2–20 mm, minutely glandular-puberulent.

Leaves

basal and cauline, basal in rosette, cauline gradually reduced distally;

petiole absent;

blade obovate to narrowly elliptic, 3–46(–70) × 1–20(–25) mm, margins entire or crenate, plane, apex rounded or obtuse, surfaces: proximals glabrate, distals glandular-pubescent and viscid.

basal and cauline or usually cauline, relatively even-sized or gradually larger distally;

petiole absent, proximal base short petiole-like, 1–5 mm;

blade ovate to broadly elliptic-ovate, (10–)15–25(–35) × 4–17 mm, margins entire, plane, apex acuminate, surfaces glabrous or adaxial minutely glandular-puberulent.

Pedicels

1–3(–6) mm in fruit.

1–1.5 mm in fruit.

Flowers

1 per node, chasmogamous.

2 per node, or 1 or 2 per node on 1 plant, chasmogamous.

Styles

eglandular-puberulent.

pubescent, at least on distal 1/2.

Corollas

pink to magenta, throat floor with 3 dark purple lines meeting abaxial lip lobes, palate ridges yellow, tube-throat 7–10 mm, limb 4–7 mm diam., not bilabiate.

magenta or rose purple, tube yellow, throat yellow, throat and distal tube red-spotted on floor, palate ridges yellow, tube-throat 13–16(–19) mm, limb 16–24 mm diam., bilabiate.

Calyces

symmetrically attached to pedicels, inflated in fruit, 5–10 mm, glandular-pubescent and viscid, lobes subequal, apex obtuse, often apiculate, ribs dark green to purplish, intercostal areas whitish.

symmetrically attached to pedicels, not inflated in fruit, 7–10 mm, glabrous or minutely stipitate-glandular, lobes unequal, apex linear-acuminate, sharp-pointed, ribs green distally, intercostal areas whitish.

Capsules

7–11(–12) mm.

10–15 mm.

Anthers

nearly exserted, glabrous.

included, glabrous or sparsely hirsutulous.

Stigmas

nearly exserted, lobes unequal, abaxial 5–7 times adaxial.

exserted, lobes subequal.

= 16.

Diplacus rattanii

Diplacus cusickii

Phenology

Flowering Apr–Jul.

Flowering May–Jul.

Habitat

Recently burned or cleared areas, sandhills, sandstone outcrops, sandy gravel and loam, decomposed granite, serpentine-derived soils, open chaparral, chaparral margins, open yellow pine-manzanita woodlands.

Slopes, canyons, washes, ditches, sand talus, diatomaceous slopes, basalt outcrops, black volcanic gravel, volcanic ash and sand, sagebrush areas.

Elevation

300–1300 m. (1000–4300 ft.)

800–1000 m. (2600–3300 ft.)

Distribution

[WildflowerSearch map]

ID; OR

[WildflowerSearch map]

[BONAP county map]

Discussion

Diplacus rattanii occurs mostly in near-coastal localities from Glenn and Lake counties south to Ventura County.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Diplacus cusickii is endemic to northern Malheur County, Oregon, and along the Snake River in Ada and Owyhee counties, Idaho. Its narrow geographic range reflects the segregation of the more widely distributed D. cusickioides.

D. M. Thompson (2005) noted that collections in northern Malheur County, Oregon, were intermediate between Mimulus cusickii and M. nanus; the type of M. cusickii is from this area and is among the narrowly endemic, supposedly putative intermediates (G. L. Nesom 2013c). These plants have abruptly and sharply acuminate leaf apices like the more widespread form traditionally identified as Diplacus cusickii; they differ in having glabrous leaf surfaces. Typical D. nanus occurs in close sympatry, without intergradation, with the populations in northern Malheur County. Because of their distinctive morphology and coherent geography, the northern Malheur County plants are reasonably recognized as a distinct species. The more widely distributed form formerly identified as D. cusickii now is identified as D. cusickioides. Populations of D. cusickii in the narrow sense along the Snake River in Ada and Owyhee counties, Idaho, may have dispersed there from the Oregon center. Some plants of D. cusickioides in the Leslie Gulch area of east-central Malheur County have somewhat reduced vestiture, approaching that of D. cusickii.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 17, p. 432.

FNA vol. 17, p. 439.

Parent taxa

Phrymaceae > Diplacus

Sibling taxa

D. angustatus, D. aridus, D. aurantiacus, D. bicolor, D. bigelovii, D. bolanderi, D. brandegeei, D. brevipes, D. calycinus, D. cascadensis, D. clevelandii, D. clivicola, D. compactus, D. congdonii, D. constrictus, D. cusickii, D. cusickioides, D. deschutesensis, D. douglasii, D. fremontii, D. grandiflorus, D. graniticola, D. jepsonii, D. johnstonii, D. kelloggii, D. layneae, D. leptaleus, D. linearis, D. longiflorus, D. mephiticus, D. mohavensis, D. nanus, D. ovatus, D. parryi, D. parviflorus, D. pictus, D. pulchellus, D. puniceus, D. pygmaeus, D. rupicola, D. rutilus, D. thompsonii, D. torreyi, D. traskiae, D. tricolor, D. vandenbergensis, D. viscidus

D. angustatus, D. aridus, D. aurantiacus, D. bicolor, D. bigelovii, D. bolanderi, D. brandegeei, D. brevipes, D. calycinus, D. cascadensis, D. clevelandii, D. clivicola, D. compactus, D. congdonii, D. constrictus, D. cusickioides, D. deschutesensis, D. douglasii, D. fremontii, D. grandiflorus, D. graniticola, D. jepsonii, D. johnstonii, D. kelloggii, D. layneae, D. leptaleus, D. linearis, D. longiflorus, D. mephiticus, D. mohavensis, D. nanus, D. ovatus, D. parryi, D. parviflorus, D. pictus, D. pulchellus, D. puniceus, D. pygmaeus, D. rattanii, D. rupicola, D. rutilus, D. thompsonii, D. torreyi, D. traskiae, D. tricolor, D. vandenbergensis, D. viscidus

Synonyms

Mimulus rattanii, M. rattanii subsp. decurtatus

Eunanus cusickii, Mimulus cusickii

Name authority

(A. Gray) G. L. Nesom: Phytoneuron 2012-39: 29. (2012)

(Greene) G. L. Nesom: Phytoneuron 2012-39: 28. (2012)

Web links

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

Choose a data source: Flora of North America, OregonFlora

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Diplacus rattanii

Diplacus cusickii

Diplacus rattanii

Diplacus cusickii