Diplacus mephiticus |
Diplacus parviflorus |
|
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foul odor monkeyflower, skunky monkeyflower |
island bush monkeyflower |
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Habit | Herbs, annual. | Subshrubs or shrubs. |
Stems | erect, (20–)30–150(–180) mm, glandular-pubescent and viscid-villous. |
erect to ascending-erect, 300–1200 mm, glabrous. |
Leaves | usually cauline, relatively even-sized; petiole absent; blade ovate to oblong or narrowly elliptic to narrowly lanceolate, narrowly oblong, or linear, 10–25 × 1–5 mm, margins entire, plane, not ciliate, apex rounded to obtuse or acute, surfaces usually glandular-pubescent and (at least along veins) viscid-villous. |
cauline, relatively even-sized; petiole absent; blade elliptic to broadly elliptic-oblanceolate or elliptic-lanceolate, 15–60 × 4–21 mm, margins entire or serrate, plane or revolute, apex acute to obtuse or rounded, surfaces glabrous. |
Pedicels | 1–3 mm in fruit. |
6–15 mm in fruit. |
Flowers | 2 per node, or 1 or 2 per node on 1 plant, chasmogamous. |
2 per node, chasmogamous. |
Styles | glabrous or sparsely glandular-puberulent. |
minutely glandular. |
Corollas | of 2 color forms: (a) dark magenta, purplish, or reddish with palate ridges or whole throat floor yellow, red- or purple-dotted, lateral lobes yellowish inside and (b) yellow with red or purple spots on floor, tube-throat 8–12(–15) mm, tube 1.3–1.9 mm diam. at filament insertion, limb 5–12(–15) mm diam., bilabiate. |
deep red to scarlet, throat sometimes orange, not spotted or striped, palate ridges red to orange-red, tube-throat 27–33 mm, limb 12–16 mm diam., bilabiate, lobes oblong, each truncate-entire to slightly emarginate. |
Calyces | symmetrically attached to pedicels, slightly inflated in fruit, (3–)4–7(–9) mm, glandular-pubescent and viscid-villous (at least along veins), lobes subequal, apex acute to acuminate, ribs dark green to purplish, intercostal areas whitish. |
not inflated in fruit, 18–25 mm, glabrous, tube slightly dilated distally, lobes unequal, apex acute, ribs green, intercostal areas light green. |
Capsules | 5–8 mm. |
14–21 mm. |
Anthers | exserted, short-hirsute. |
exserted, glabrous. |
Stigmas | exserted or at opening of corolla tube-throat, lobes subequal to unequal, abaxial to 2 times adaxial. |
exserted, lobes equal. |
2n | = 20. |
|
Diplacus mephiticus |
Diplacus parviflorus |
|
Phenology | Flowering (May–)Jun–Aug(–Sep). | Flowering Mar–Aug. |
Habitat | Openings in sagebrush, disturbed slopes, granite outcrops, serpentine substrates, gravelly and sandy soils, sandy moraines, pumice flats, gravelly washes, meadows, shadscale and sagebrush communities, pinyon-juniper and ponderosa pine woodlands. | Hillsides, canyons, rocky slopes and walls, bluffs, sea cliffs. |
Elevation | 1300–3700 m. (4300–12100 ft.) | 10–400 m. (0–1300 ft.) |
Distribution |
CA; NV
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CA
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Discussion | Diplacus mephiticus occurs in eastern California and west-central Nevada. Various synonyms treated here are in agreement with D. M. Thompson (2005). Mimulus coccineus (mostly from Eldorado to Tulare counties, California, and, apparently, including Eunanus angustifolius Greene from Mt. Rose, Nevada) includes relatively small, tufted plants at high elevations with relatively small calyces and relatively small, dark red-purple, strongly bilabiate corollas with prominently exserted stamens. Mimulus densus (mostly in Nevada and in Lassen, Nevada, and Plumas counties, California) includes taller plants at lower elevations with a strong tendency to produce populations with all individuals with larger, yellow, nearly regular corollas with more nearly included stamens. Typical Diplacus mephiticus has moderate-sized plants at medium elevations with magenta, bilabiate corollas. The specific epithet mephiticus alludes to the musky odor of the plants; this has also been noted in plants of Mimulus coccineus and M. densus. Diplacus nanus, in which D. M. Thompson (2005) included D. mephiticus as a variety, apparently does not produce a mephitic odor. Diplacus cusickii also produces a mephitic odor (W. L. Ezell 1971). The later homonym Mimulus angustifolius (Greene) A. L. Grant 1925, not Hochstetter ex Richard 1850, based on Eunanus angustifolius, pertains here. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Diplacus parviflorus is known from four of the Channel Islands (Anacapa, San Clemente, Santa Cruz, and Santa Rosa). Hybrids with Diplacus longiflorus occur on Santa Cruz Island; A. L. Grant (according to label data of collections) found these to be fairly common on open hillsides near Friar’s Harbor and Valdez, where the two species grew near each other though apparently separated in habitat, with typical D. parviflorus mostly in the canyons and D. longiflorus on open hillsides. She noted that the apparent hybrids were variable in all possible combinations of features of the leaves, calyces, and corollas, including color. Mimulus parviflorus (Greene) A. L. Grant 1925, not Lindley 1825, pertains here. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 441. | FNA vol. 17, p. 452. |
Parent taxa | Phrymaceae > Diplacus | Phrymaceae > Diplacus |
Sibling taxa | ||
Synonyms | Mimulus mephiticus, M. coccineus, M. coccineus var. wolfii, M. densus, M. nanus var. mephiticus, M. reifschneiderae, M. stamineus, M. washoensis, M. wolfii | Mimulus aurantiacus var. parviflorus, M. flemingii |
Name authority | (Greene) G. L. Nesom: Phytoneuron 2012-39: 29. (2012) | Greene: Pittonia 1: 36. (1887) |
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