Diplacus mephiticus |
Diplacus johnstonii |
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foul odor monkeyflower, skunky monkeyflower |
Johnston's monkeyflower |
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Habit | Herbs, annual. | Herbs, annual. |
Stems | erect, (20–)30–150(–180) mm, glandular-pubescent and viscid-villous. |
erect, (10–)30–200(–300) mm, densely glandular-puberulent. |
Leaves | usually cauline, relatively even-sized; petiole absent; blade ovate to oblong or narrowly elliptic to narrowly lanceolate, narrowly oblong, or linear, 10–25 × 1–5 mm, margins entire, plane, not ciliate, apex rounded to obtuse or acute, surfaces usually glandular-pubescent and (at least along veins) viscid-villous. |
basal and cauline, gradually reduced distally; petiole absent, bases of larger leaves often with petiole-like extensions; blade obovate or oblanceolate, sometimes elliptic, (4.5–)7–25(–32) × 2–12(–15) mm, margins entire, plane, apex rounded to acute, surfaces densely glandular-puberulent. |
Pedicels | 1–3 mm in fruit. |
1–4(–5) mm in fruit. |
Flowers | 2 per node, or 1 or 2 per node on 1 plant, chasmogamous. |
2 per node, or 1 or 2 per node on 1 plant, chasmogamous. |
Styles | glabrous or sparsely glandular-puberulent. |
densely glandular-puberulent distally. |
Corollas | of 2 color forms: (a) dark magenta, purplish, or reddish with palate ridges or whole throat floor yellow, red- or purple-dotted, lateral lobes yellowish inside and (b) yellow with red or purple spots on floor, tube-throat 8–12(–15) mm, tube 1.3–1.9 mm diam. at filament insertion, limb 5–12(–15) mm diam., bilabiate. |
magenta, darker and more reddish in throat and, often, along narrow radiating lines extending from throat onto midveins of lobes, throat usually with a large dark spot on each side of mouth on lateral walls, palate ridges and throat floor yellow with reddish spots, tube-throat 9–15 mm, limb 10–15 mm diam., not bilabiate. |
Calyces | symmetrically attached to pedicels, slightly inflated in fruit, (3–)4–7(–9) mm, glandular-pubescent and viscid-villous (at least along veins), lobes subequal, apex acute to acuminate, ribs dark green to purplish, intercostal areas whitish. |
symmetrically attached to pedicels, not inflated in fruit, (6–)7–11 mm, glandular-puberulent, lobes unequal, apex acute to acuminate, ribs inconspicuous, intercostal areas reddish. |
Capsules | 5–8 mm. |
7–12 mm. |
Anthers | exserted, short-hirsute. |
included, glabrous. |
Stigmas | exserted or at opening of corolla tube-throat, lobes subequal to unequal, abaxial to 2 times adaxial. |
exserted or at opening of corolla tube-throat, lobes equal. |
2n | = 16. |
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Diplacus mephiticus |
Diplacus johnstonii |
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Phenology | Flowering (May–)Jun–Aug(–Sep). | Flowering May–Aug. |
Habitat | Openings in sagebrush, disturbed slopes, granite outcrops, serpentine substrates, gravelly and sandy soils, sandy moraines, pumice flats, gravelly washes, meadows, shadscale and sagebrush communities, pinyon-juniper and ponderosa pine woodlands. | Steep, unstable scree slides, talus slopes, gravel slides, cracks in granite cliffs, ridges, washes, steep sand and gravel slopes, canyon bottoms, gravelly road banks, recent burns, desert scrub, chaparral, juniper, pinyon-juniper, lodgepole pine, yellow pine, Jeffrey pine, and Jeffrey pine-western white pine-fir woodlands. |
Elevation | 1300–3700 m. (4300–12100 ft.) | (1000–)1300–2900 m. ((3300–)4300–9500 ft.) |
Distribution |
CA; NV
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CA
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Discussion | Diplacus mephiticus occurs in eastern California and west-central Nevada. Various synonyms treated here are in agreement with D. M. Thompson (2005). Mimulus coccineus (mostly from Eldorado to Tulare counties, California, and, apparently, including Eunanus angustifolius Greene from Mt. Rose, Nevada) includes relatively small, tufted plants at high elevations with relatively small calyces and relatively small, dark red-purple, strongly bilabiate corollas with prominently exserted stamens. Mimulus densus (mostly in Nevada and in Lassen, Nevada, and Plumas counties, California) includes taller plants at lower elevations with a strong tendency to produce populations with all individuals with larger, yellow, nearly regular corollas with more nearly included stamens. Typical Diplacus mephiticus has moderate-sized plants at medium elevations with magenta, bilabiate corollas. The specific epithet mephiticus alludes to the musky odor of the plants; this has also been noted in plants of Mimulus coccineus and M. densus. Diplacus nanus, in which D. M. Thompson (2005) included D. mephiticus as a variety, apparently does not produce a mephitic odor. Diplacus cusickii also produces a mephitic odor (W. L. Ezell 1971). The later homonym Mimulus angustifolius (Greene) A. L. Grant 1925, not Hochstetter ex Richard 1850, based on Eunanus angustifolius, pertains here. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Populations of Diplacus johnstonii occur in Los Angeles, Riverside, San Bernardino, Santa Barbara, and Ventura counties. The populations in northwestern Los Angeles, Santa Barbara, and Ventura counties were noted by D. M. Thompson (2005) to be intermediate between D. constrictus and D. johnstonii. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 441. | FNA vol. 17, p. 433. |
Parent taxa | Phrymaceae > Diplacus | Phrymaceae > Diplacus |
Sibling taxa | ||
Synonyms | Mimulus mephiticus, M. coccineus, M. coccineus var. wolfii, M. densus, M. nanus var. mephiticus, M. reifschneiderae, M. stamineus, M. washoensis, M. wolfii | Mimulus johnstonii |
Name authority | (Greene) G. L. Nesom: Phytoneuron 2012-39: 29. (2012) | (A. L. Grant) G. L. Nesom: Phytoneuron 2012-39: 29. (2012) |
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