Diplacus mephiticus |
Diplacus angustatus |
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foul odor monkeyflower, skunky monkeyflower |
narrow leaf pansy monkeyflower, narrow-leaf or purple-lip pansy monkeyflower, purplelip pansy monkeyflower |
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Habit | Herbs, annual. | Herbs, annual, acaulescent or caulescent. |
Stems | erect, (20–)30–150(–180) mm, glandular-pubescent and viscid-villous. |
erect, 0 or 20–70 mm, eglandular-puberulent. |
Leaves | usually cauline, relatively even-sized; petiole absent; blade ovate to oblong or narrowly elliptic to narrowly lanceolate, narrowly oblong, or linear, 10–25 × 1–5 mm, margins entire, plane, not ciliate, apex rounded to obtuse or acute, surfaces usually glandular-pubescent and (at least along veins) viscid-villous. |
basal densely clustered; petiole absent; blade narrowly lanceolate or elliptic to linear, (3.5–)5–36 × 0.5–3.8(–5) mm, margins entire, plane, proximal 1/2+ ciliate, apex obtuse, surfaces glabrous. |
Pedicels | 1–3 mm in fruit. |
0–1 mm in fruit. |
Flowers | 2 per node, or 1 or 2 per node on 1 plant, chasmogamous. |
1 per node, chasmogamous. |
Styles | glabrous or sparsely glandular-puberulent. |
glandular-puberulent. |
Corollas | of 2 color forms: (a) dark magenta, purplish, or reddish with palate ridges or whole throat floor yellow, red- or purple-dotted, lateral lobes yellowish inside and (b) yellow with red or purple spots on floor, tube-throat 8–12(–15) mm, tube 1.3–1.9 mm diam. at filament insertion, limb 5–12(–15) mm diam., bilabiate. |
throat and limb magenta to purple with a dark maroon-purple spot or elongate blotch at base of each abaxial lobe, palate ridges and base of abaxial lip yellow, tube-throat 20–60 mm, limb 10–20 mm diam., bilabiate, lobes equal. |
Calyces | symmetrically attached to pedicels, slightly inflated in fruit, (3–)4–7(–9) mm, glandular-pubescent and viscid-villous (at least along veins), lobes subequal, apex acute to acuminate, ribs dark green to purplish, intercostal areas whitish. |
slightly asymmetrically attached to pedicel, inflated in fruit, (4–)7–14(–17) mm, pilose, lobes subequal, apex obtuse, ribs becoming reddish, intercostal areas pale green to whitish. |
Capsules | 5–8 mm. |
2–4.5 mm, indehiscent. |
Anthers | exserted, short-hirsute. |
(distal pair) sometimes nearly exserted, hirsute with 2 apical tufts. |
Stigmas | exserted or at opening of corolla tube-throat, lobes subequal to unequal, abaxial to 2 times adaxial. |
usually exserted, lobes subequal. |
2n | = 18. |
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Diplacus mephiticus |
Diplacus angustatus |
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Phenology | Flowering (May–)Jun–Aug(–Sep). | Flowering Mar–May(–Jun). |
Habitat | Openings in sagebrush, disturbed slopes, granite outcrops, serpentine substrates, gravelly and sandy soils, sandy moraines, pumice flats, gravelly washes, meadows, shadscale and sagebrush communities, pinyon-juniper and ponderosa pine woodlands. | Vernally flooded depressions or swales, summits of mesas or foothills. |
Elevation | 1300–3700 m. (4300–12100 ft.) | 300–1200 m. (1000–3900 ft.) |
Distribution |
CA; NV
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CA
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Discussion | Diplacus mephiticus occurs in eastern California and west-central Nevada. Various synonyms treated here are in agreement with D. M. Thompson (2005). Mimulus coccineus (mostly from Eldorado to Tulare counties, California, and, apparently, including Eunanus angustifolius Greene from Mt. Rose, Nevada) includes relatively small, tufted plants at high elevations with relatively small calyces and relatively small, dark red-purple, strongly bilabiate corollas with prominently exserted stamens. Mimulus densus (mostly in Nevada and in Lassen, Nevada, and Plumas counties, California) includes taller plants at lower elevations with a strong tendency to produce populations with all individuals with larger, yellow, nearly regular corollas with more nearly included stamens. Typical Diplacus mephiticus has moderate-sized plants at medium elevations with magenta, bilabiate corollas. The specific epithet mephiticus alludes to the musky odor of the plants; this has also been noted in plants of Mimulus coccineus and M. densus. Diplacus nanus, in which D. M. Thompson (2005) included D. mephiticus as a variety, apparently does not produce a mephitic odor. Diplacus cusickii also produces a mephitic odor (W. L. Ezell 1971). The later homonym Mimulus angustifolius (Greene) A. L. Grant 1925, not Hochstetter ex Richard 1850, based on Eunanus angustifolius, pertains here. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Diplacus angustatus occurs in northern California in the Sierra Nevada and across the Sacramento Valley in the North Coast Range (Lake, Mendocino, and Napa counties); it also is known from one area in northwestern Shasta County and has a disjunct series of populations in Fresno and Tulare counties. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 441. | FNA vol. 17, p. 447. |
Parent taxa | Phrymaceae > Diplacus | Phrymaceae > Diplacus |
Sibling taxa | ||
Synonyms | Mimulus mephiticus, M. coccineus, M. coccineus var. wolfii, M. densus, M. nanus var. mephiticus, M. reifschneiderae, M. stamineus, M. washoensis, M. wolfii | Eunanus coulteri var. angustatus, E. angustatus, Mimulus angustatus, M. tricolor var. angustatus |
Name authority | (Greene) G. L. Nesom: Phytoneuron 2012-39: 29. (2012) | (A. Gray) G. L. Nesom: Phytoneuron 2012-39: 30. (2012) |
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