Flora of North America species comparison

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Diplacus layneae

Diplacus vandenbergensis

Layne's monkeyflower

Vandenberg monkeyflower
Habit Herbs, annual, herbage usually drying dark. Herbs, annual.
Stems

erect, 30–160(–300) mm, nodes 3–6, glandular-puberulent to glandular-pubescent, hairs 0.2–0.8 mm.

erect, 10–200(–240) mm, glandular-puberulent or glandular-pubescent.
Leaves

usually cauline, relatively even-sized;

petiole weakly delimited;

blade elliptic to narrowly elliptic, oblanceolate, elliptic-oblanceolate, or elliptic-lanceolate, 8–27(–35) × 2–8 mm, margins entire, rarely toothed, plane, apex rounded to obtuse, surfaces: proximals often glabrate, distals glandular-puberulent or glandular-pubescent.

basal and cauline, basal in rosette, cauline reduced distally;

petiole absent;

blade narrowly elliptic, sometimes obovate to oblanceolate, 2–30(–55) × 1–10(–16) mm, margins entire, sometimes crenate to serrate, plane, not ciliate, apex rounded to acute, surfaces: proximals glabrous, distals glandular-puberulent or glandular-pubescent.
Pedicels

2–4(–5) mm in fruit.

1–4(–7 on proximal) mm in fruit.
Flowers

2 per node, or 1 or 2 per node on 1 plant, chasmogamous.

1 per node, chasmogamous.
Styles

glandular-puberulent.

minutely puberulent.
Corollas

pinkish or pale to dark magenta or red-purple, each lobe usually with a faint to dark medial line extending 1/2 or less to tip, throat floor yellowish near base, mostly white with red-purple dots near mouth, palate ridges white, tube-throat 10–15 mm, limb (8–)10–16 mm diam., not bilabiate.

yellow with reddish brown spots near mouth, palate ridges yellow, throat floor puberulent, tube-throat (9–)10–23 mm, limb 8–26 mm diam., not bilabiate.
Calyces

symmetrically attached to pedicels, not inflated in fruit, (5–)6–8(–9) mm, glandular-puberulent to glandular-pubescent, tube strongly plicate, lobes triangular, subequal, apex acute, ribs broad, darkened, blackish, thickened, strongly raised, intercostal areas whitish, membranous.

inflated in fruit, 5–14 mm, glandular-puberulent to glandular-pubescent or ribs almost tomentose and viscid, lobes subequal, apex rounded and apiculate or acute, intercostal areas white.
Capsules

6–10(–13) mm.

6.5–13(–14) mm.
Anthers

included, ciliate.

included, glabrous, rarely minutely puberulent.
Stigmas

included, lobes unequal, abaxial 1.5 times adaxial.

included, lobes equal.
2n

= 16.

= 16.

Diplacus layneae

Diplacus vandenbergensis
Phenology Flowering May–Aug. Flowering Apr–Jun.
Habitat Road banks, serpentine, granitic sand, red clay, lava beds and volcanic soils, openings in chaparral, shallow dry streambeds or stream banks, burned or otherwise disturbed open areas. Sandy open or disturbed areas, among shrubs.
Elevation (100–)400–2400 m. ((300–)1300–7900 ft.) 70–150 m. (200–500 ft.)
Distribution
from FNA
CA
[WildflowerSearch map]
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
Discussion

D. M. Thompson (2005) observed that two forms of Diplacus layneae co-occur from the Yosemite National Park area southward; one of these is recognized here as D. graniticola.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Diplacus vandenbergensis is endemic to Santa Barbara County, mostly on the north side of the city of Lompoc (La Purisima Mission State Historic Park; also on and near Vandenberg Air Force Base) plus one other locality about 10 kilometers farther west (Santa Ynez Valley, 8 miles west of Buellton). Thompson described Mimulus fremontii var. vandenbergensis as a geographically distinct variant of M. fremontii. The discontinuous morphological difference, allopatric/parapatric geographical distribution, and lack of intergrading populations support recognition of the plants with yellow corollas at specific rank. Plants with yellow corollas are found in D. bicolor, D. mephiticus, and D. parryi, but these morphs are found through most of the geographic ranges of these taxa and sometimes are intermixed in populations.

(Discussion copyrighted by Flora of North America; reprinted with permission.)
Source FNA vol. 17, p. 437. FNA vol. 17, p. 432.
Parent taxa Phrymaceae > Diplacus Phrymaceae > Diplacus
Sibling taxa
D. angustatus, D. aridus, D. aurantiacus, D. bicolor, D. bigelovii, D. bolanderi, D. brandegeei, D. brevipes, D. calycinus, D. cascadensis, D. clevelandii, D. clivicola, D. compactus, D. congdonii, D. constrictus, D. cusickii, D. cusickioides, D. deschutesensis, D. douglasii, D. fremontii, D. grandiflorus, D. graniticola, D. jepsonii, D. johnstonii, D. kelloggii, D. leptaleus, D. linearis, D. longiflorus, D. mephiticus, D. mohavensis, D. nanus, D. ovatus, D. parryi, D. parviflorus, D. pictus, D. pulchellus, D. puniceus, D. pygmaeus, D. rattanii, D. rupicola, D. rutilus, D. thompsonii, D. torreyi, D. traskiae, D. tricolor, D. vandenbergensis, D. viscidus
D. angustatus, D. aridus, D. aurantiacus, D. bicolor, D. bigelovii, D. bolanderi, D. brandegeei, D. brevipes, D. calycinus, D. cascadensis, D. clevelandii, D. clivicola, D. compactus, D. congdonii, D. constrictus, D. cusickii, D. cusickioides, D. deschutesensis, D. douglasii, D. fremontii, D. grandiflorus, D. graniticola, D. jepsonii, D. johnstonii, D. kelloggii, D. layneae, D. leptaleus, D. linearis, D. longiflorus, D. mephiticus, D. mohavensis, D. nanus, D. ovatus, D. parryi, D. parviflorus, D. pictus, D. pulchellus, D. puniceus, D. pygmaeus, D. rattanii, D. rupicola, D. rutilus, D. thompsonii, D. torreyi, D. traskiae, D. tricolor, D. viscidus
Synonyms Eunanus layneae, Mimulus brachiatus, M. layneae Mimulus fremontii var. vandenbergensis
Name authority (Greene) G. L. Nesom: Phytoneuron 2012-39: 29. (2012) (D. M. Thompson) G. L. Nesom: Phytoneuron 2012-47: 2. (2012)
Web links 
Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).



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