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Layne's monkeyflower

Death Valley monkeyflower, rock midget

Habit Herbs, annual, herbage usually drying dark. Herbs, perennial, with woody caudex.
Stems

erect, 30–160(–300) mm, nodes 3–6, glandular-puberulent to glandular-pubescent, hairs 0.2–0.8 mm.

erect to ascending, sometimes pendent, 10–170 mm, densely and finely glandular-puberulent.

Leaves

usually cauline, relatively even-sized;

petiole weakly delimited;

blade elliptic to narrowly elliptic, oblanceolate, elliptic-oblanceolate, or elliptic-lanceolate, 8–27(–35) × 2–8 mm, margins entire, rarely toothed, plane, apex rounded to obtuse, surfaces: proximals often glabrate, distals glandular-puberulent or glandular-pubescent.

usually basal rosettes and proximal cauline, relatively even-sized;

petiole absent, base gradually narrowed to broad, petiole-like extension;

blade oblanceolate, (10–)18–60(–80) × (1.5–)3–15(–26) mm, margins entire, plane, not ciliate, apex acute, surfaces glandular-puberulent.

Pedicels

2–4(–5) mm in fruit.

1–3 mm in fruit.

Flowers

2 per node, or 1 or 2 per node on 1 plant, chasmogamous.

1 or 2 per node, chasmogamous.

Styles

glandular-puberulent.

glandular-puberulent.

Corollas

pinkish or pale to dark magenta or red-purple, each lobe usually with a faint to dark medial line extending 1/2 or less to tip, throat floor yellowish near base, mostly white with red-purple dots near mouth, palate ridges white, tube-throat 10–15 mm, limb (8–)10–16 mm diam., not bilabiate.

limb pinkish white to nearly white with a large magenta-purple round or 2-lobed blotch at base of each lobe, throat and palate ridges golden yellow with magenta speckling, palate ridges short-pilose, throat glabrous, tube-throat 17–35 mm, limb 8–21 mm diam., not bilabiate.

Calyces

symmetrically attached to pedicels, not inflated in fruit, (5–)6–8(–9) mm, glandular-puberulent to glandular-pubescent, tube strongly plicate, lobes triangular, subequal, apex acute, ribs broad, darkened, blackish, thickened, strongly raised, intercostal areas whitish, membranous.

asymmetrically attached to pedicel, not inflated in fruit, 8–18 mm, densely glandular-puberulent, lobes unequal, apex acuminate, ribs green, intercostal areas pale green.

Capsules

6–10(–13) mm.

3–8 mm, indehiscent until senescence of pedicel, then opening along both sutures only after wetting.

Anthers

included, ciliate.

included, glabrous or slightly puberulent at base.

Stigmas

included, lobes unequal, abaxial 1.5 times adaxial.

included, lobes equal.

2n

= 16.

= 16.

Diplacus layneae

Diplacus rupicola

Phenology Flowering May–Aug. Flowering Feb–Jun.
Habitat Road banks, serpentine, granitic sand, red clay, lava beds and volcanic soils, openings in chaparral, shallow dry streambeds or stream banks, burned or otherwise disturbed open areas. Crevices in limestone cliffs and walls, limestone ridge tops and slopes, wash edges, gravelly slopes, canyon sides.
Elevation (100–)400–2400 m. ((300–)1300–7900 ft.) 300–1800 m. (1000–5900 ft.)
Distribution
from FNA
CA
[WildflowerSearch map]
from FNA
CA
[WildflowerSearch map]
Discussion

D. M. Thompson (2005) observed that two forms of Diplacus layneae co-occur from the Yosemite National Park area southward; one of these is recognized here as D. graniticola.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Diplacus rupicola is known from Inyo County.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 437. FNA vol. 17, p. 448.
Parent taxa Phrymaceae > Diplacus Phrymaceae > Diplacus
Sibling taxa
D. angustatus, D. aridus, D. aurantiacus, D. bicolor, D. bigelovii, D. bolanderi, D. brandegeei, D. brevipes, D. calycinus, D. cascadensis, D. clevelandii, D. clivicola, D. compactus, D. congdonii, D. constrictus, D. cusickii, D. cusickioides, D. deschutesensis, D. douglasii, D. fremontii, D. grandiflorus, D. graniticola, D. jepsonii, D. johnstonii, D. kelloggii, D. leptaleus, D. linearis, D. longiflorus, D. mephiticus, D. mohavensis, D. nanus, D. ovatus, D. parryi, D. parviflorus, D. pictus, D. pulchellus, D. puniceus, D. pygmaeus, D. rattanii, D. rupicola, D. rutilus, D. thompsonii, D. torreyi, D. traskiae, D. tricolor, D. vandenbergensis, D. viscidus
D. angustatus, D. aridus, D. aurantiacus, D. bicolor, D. bigelovii, D. bolanderi, D. brandegeei, D. brevipes, D. calycinus, D. cascadensis, D. clevelandii, D. clivicola, D. compactus, D. congdonii, D. constrictus, D. cusickii, D. cusickioides, D. deschutesensis, D. douglasii, D. fremontii, D. grandiflorus, D. graniticola, D. jepsonii, D. johnstonii, D. kelloggii, D. layneae, D. leptaleus, D. linearis, D. longiflorus, D. mephiticus, D. mohavensis, D. nanus, D. ovatus, D. parryi, D. parviflorus, D. pictus, D. pulchellus, D. puniceus, D. pygmaeus, D. rattanii, D. rutilus, D. thompsonii, D. torreyi, D. traskiae, D. tricolor, D. vandenbergensis, D. viscidus
Synonyms Eunanus layneae, Mimulus brachiatus, M. layneae Mimulus rupicola
Name authority (Greene) G. L. Nesom: Phytoneuron 2012-39: 29. (2012) (Coville & A. L. Grant) G. L. Nesom & N. S. Fraga: Phytoneuron 2012-39: 27. (2012)
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