FNA: Diplacus layneae vs. Diplacus jepsonii

	Diplacus layneae	Diplacus jepsonii
	Layne's monkeyflower	Jepson's monkeyflower
Habit	Herbs, annual, herbage usually drying dark.	Herbs, annual.
Stems	erect, 30–160(–300) mm, nodes 3–6, glandular-puberulent to glandular-pubescent, hairs 0.2–0.8 mm.	erect, 10–150 mm, minutely glandular-puberulent.
Leaves	usually cauline, relatively even-sized; petiole weakly delimited; blade elliptic to narrowly elliptic, oblanceolate, elliptic-oblanceolate, or elliptic-lanceolate, 8–27(–35) × 2–8 mm, margins entire, rarely toothed, plane, apex rounded to obtuse, surfaces: proximals often glabrate, distals glandular-puberulent or glandular-pubescent.	usually cauline, relatively even-sized; petiole absent; blade narrowly elliptic-oblanceolate to narrowly oblanceolate, 6–22 × 1–5(–7) mm, margins entire, plane, apex acute to obtuse, surfaces glandular-puberulent to glandular-pubescent.
Pedicels	2–4(–5) mm in fruit.	1 mm in fruit.
Flowers	2 per node, or 1 or 2 per node on 1 plant, chasmogamous.	2 per node, or 1 or 2 per node on 1 plant, chasmogamous.
Styles	glandular-puberulent.	glandular-puberulent or glandular-pubescent.
Corollas	pinkish or pale to dark magenta or red-purple, each lobe usually with a faint to dark medial line extending 1/2 or less to tip, throat floor yellowish near base, mostly white with red-purple dots near mouth, palate ridges white, tube-throat 10–15 mm, limb (8–)10–16 mm diam., not bilabiate.	lavender-purple to rose purple, throat darker, palate ridges yellow, red-spotted, tube 0.8–1.2 mm diam. at filament insertion, tube-throat 7–10 mm, limb 5–7 mm diam., bilabiate.
Calyces	symmetrically attached to pedicels, not inflated in fruit, (5–)6–8(–9) mm, glandular-puberulent to glandular-pubescent, tube strongly plicate, lobes triangular, subequal, apex acute, ribs broad, darkened, blackish, thickened, strongly raised, intercostal areas whitish, membranous.	symmetrically attached to pedicels, not inflated in fruit, 3–5 mm, minutely puberulent, lobes subequal, apex acute to acuminate, ribs dark green to purplish, intercostal areas whitish.
Capsules	6–10(–13) mm.	4–6 mm.
Anthers	included, ciliate.	exserted, sparsely hispidulous.
Stigmas	included, lobes unequal, abaxial 1.5 times adaxial.	exserted, lobes equal.
2n	= 16.	= 16.

	Diplacus layneae	Diplacus jepsonii
Phenology	Flowering May–Aug.	Flowering Jun–Aug.
Habitat	Road banks, serpentine, granitic sand, red clay, lava beds and volcanic soils, openings in chaparral, shallow dry streambeds or stream banks, burned or otherwise disturbed open areas.	Gentle slopes, sandy and volcanic soils, meadows in spruce-fir forests, openings among pines and in chaparral, shallow drainage areas, disturbed open areas.
Elevation	(100–)400–2400 m. ((300–)1300–7900 ft.)	(900–)1100–2500(–2800) m. ((3000–)3600–8200(–9200) ft.)
Distribution	CA [WildflowerSearch map]	CA; NV; OR [WildflowerSearch map]
Discussion	D. M. Thompson (2005) observed that two forms of Diplacus layneae co-occur from the Yosemite National Park area southward; one of these is recognized here as D. graniticola. (Discussion copyrighted by Flora of North America; reprinted with permission.)	D. M. Thompson (2005) treated Diplacus jepsonii as conspecific with D. nanus [as W. L. Ezell (1971) had done earlier] because of putative intergrades. In its typical form, D. jepsonii is immediately distinct in its narrower leaves, stems with well-separated proximal nodes, and smaller corollas with nearly filiform tubes. R. J. Meinke (1992), in a field and greenhouse study not cited by Thompson, found that D. jepsonii and D. nanus are distinct in morphology as well as in habitat and that the two do not grow intermixed in nature. Diplacus jepsonii is known only from north-central California, southern Washoe County, Nevada, and southern Oregon. Putative outliers in the central Sierra Nevada are dubiously identified. (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 17, p. 437.	FNA vol. 17, p. 442.
Parent taxa	Phrymaceae > Diplacus	Phrymaceae > Diplacus
Sibling taxa	D. angustatus, D. aridus, D. aurantiacus, D. bicolor, D. bigelovii, D. bolanderi, D. brandegeei, D. brevipes, D. calycinus, D. cascadensis, D. clevelandii, D. clivicola, D. compactus, D. congdonii, D. constrictus, D. cusickii, D. cusickioides, D. deschutesensis, D. douglasii, D. fremontii, D. grandiflorus, D. graniticola, D. jepsonii, D. johnstonii, D. kelloggii, D. leptaleus, D. linearis, D. longiflorus, D. mephiticus, D. mohavensis, D. nanus, D. ovatus, D. parryi, D. parviflorus, D. pictus, D. pulchellus, D. puniceus, D. pygmaeus, D. rattanii, D. rupicola, D. rutilus, D. thompsonii, D. torreyi, D. traskiae, D. tricolor, D. vandenbergensis, D. viscidus	D. angustatus, D. aridus, D. aurantiacus, D. bicolor, D. bigelovii, D. bolanderi, D. brandegeei, D. brevipes, D. calycinus, D. cascadensis, D. clevelandii, D. clivicola, D. compactus, D. congdonii, D. constrictus, D. cusickii, D. cusickioides, D. deschutesensis, D. douglasii, D. fremontii, D. grandiflorus, D. graniticola, D. johnstonii, D. kelloggii, D. layneae, D. leptaleus, D. linearis, D. longiflorus, D. mephiticus, D. mohavensis, D. nanus, D. ovatus, D. parryi, D. parviflorus, D. pictus, D. pulchellus, D. puniceus, D. pygmaeus, D. rattanii, D. rupicola, D. rutilus, D. thompsonii, D. torreyi, D. traskiae, D. tricolor, D. vandenbergensis, D. viscidus
Synonyms	Eunanus layneae, Mimulus brachiatus, M. layneae	Mimulus jepsonii, M. microcarpus, M. nanus var. jepsonii
Name authority	(Greene) G. L. Nesom: Phytoneuron 2012-39: 29. (2012)	(A. L. Grant) G. L. Nesom: Phytoneuron 2012-39: 29. (2012)
Web links	Local floras: CA Local Web sites: CalFlora, CalPhotos WildflowerSearch iNaturalist (observations) LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	Local floras: CA, OR Local Web sites: CalFlora, CalPhotos, Flora NW, PNW Herbaria WildflowerSearch iNaturalist (observations) LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Diplacus jepsonii

Layne's monkeyflower

Jepson's monkeyflower

Habit

Herbs, annual, herbage usually drying dark.

Herbs, annual.

Stems

erect, 30–160(–300) mm, nodes 3–6, glandular-puberulent to glandular-pubescent, hairs 0.2–0.8 mm.

erect, 10–150 mm, minutely glandular-puberulent.

Leaves

usually cauline, relatively even-sized;

petiole weakly delimited;

blade elliptic to narrowly elliptic, oblanceolate, elliptic-oblanceolate, or elliptic-lanceolate, 8–27(–35) × 2–8 mm, margins entire, rarely toothed, plane, apex rounded to obtuse, surfaces: proximals often glabrate, distals glandular-puberulent or glandular-pubescent.

usually cauline, relatively even-sized;

petiole absent;

blade narrowly elliptic-oblanceolate to narrowly oblanceolate, 6–22 × 1–5(–7) mm, margins entire, plane, apex acute to obtuse, surfaces glandular-puberulent to glandular-pubescent.

Pedicels

2–4(–5) mm in fruit.

1 mm in fruit.

Flowers

2 per node, or 1 or 2 per node on 1 plant, chasmogamous.

Styles

glandular-puberulent.

glandular-puberulent or glandular-pubescent.

Corollas

pinkish or pale to dark magenta or red-purple, each lobe usually with a faint to dark medial line extending 1/2 or less to tip, throat floor yellowish near base, mostly white with red-purple dots near mouth, palate ridges white, tube-throat 10–15 mm, limb (8–)10–16 mm diam., not bilabiate.

lavender-purple to rose purple, throat darker, palate ridges yellow, red-spotted, tube 0.8–1.2 mm diam. at filament insertion, tube-throat 7–10 mm, limb 5–7 mm diam., bilabiate.

Calyces

symmetrically attached to pedicels, not inflated in fruit, (5–)6–8(–9) mm, glandular-puberulent to glandular-pubescent, tube strongly plicate, lobes triangular, subequal, apex acute, ribs broad, darkened, blackish, thickened, strongly raised, intercostal areas whitish, membranous.

symmetrically attached to pedicels, not inflated in fruit, 3–5 mm, minutely puberulent, lobes subequal, apex acute to acuminate, ribs dark green to purplish, intercostal areas whitish.

Capsules

6–10(–13) mm.

4–6 mm.

Anthers

included, ciliate.

exserted, sparsely hispidulous.

Stigmas

included, lobes unequal, abaxial 1.5 times adaxial.

exserted, lobes equal.

= 16.

Diplacus layneae

Diplacus jepsonii

Phenology

Flowering May–Aug.

Flowering Jun–Aug.

Habitat

Road banks, serpentine, granitic sand, red clay, lava beds and volcanic soils, openings in chaparral, shallow dry streambeds or stream banks, burned or otherwise disturbed open areas.

Gentle slopes, sandy and volcanic soils, meadows in spruce-fir forests, openings among pines and in chaparral, shallow drainage areas, disturbed open areas.

Elevation

(100–)400–2400 m. ((300–)1300–7900 ft.)

(900–)1100–2500(–2800) m. ((3000–)3600–8200(–9200) ft.)

Distribution

[WildflowerSearch map]

CA; NV; OR

[WildflowerSearch map]

Discussion

D. M. Thompson (2005) observed that two forms of Diplacus layneae co-occur from the Yosemite National Park area southward; one of these is recognized here as D. graniticola.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

D. M. Thompson (2005) treated Diplacus jepsonii as conspecific with D. nanus [as W. L. Ezell (1971) had done earlier] because of putative intergrades. In its typical form, D. jepsonii is immediately distinct in its narrower leaves, stems with well-separated proximal nodes, and smaller corollas with nearly filiform tubes. R. J. Meinke (1992), in a field and greenhouse study not cited by Thompson, found that D. jepsonii and D. nanus are distinct in morphology as well as in habitat and that the two do not grow intermixed in nature.

Diplacus jepsonii is known only from north-central California, southern Washoe County, Nevada, and southern Oregon. Putative outliers in the central Sierra Nevada are dubiously identified.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 17, p. 437.

FNA vol. 17, p. 442.

Parent taxa

Phrymaceae > Diplacus

Sibling taxa

D. angustatus, D. aridus, D. aurantiacus, D. bicolor, D. bigelovii, D. bolanderi, D. brandegeei, D. brevipes, D. calycinus, D. cascadensis, D. clevelandii, D. clivicola, D. compactus, D. congdonii, D. constrictus, D. cusickii, D. cusickioides, D. deschutesensis, D. douglasii, D. fremontii, D. grandiflorus, D. graniticola, D. jepsonii, D. johnstonii, D. kelloggii, D. leptaleus, D. linearis, D. longiflorus, D. mephiticus, D. mohavensis, D. nanus, D. ovatus, D. parryi, D. parviflorus, D. pictus, D. pulchellus, D. puniceus, D. pygmaeus, D. rattanii, D. rupicola, D. rutilus, D. thompsonii, D. torreyi, D. traskiae, D. tricolor, D. vandenbergensis, D. viscidus

D. angustatus, D. aridus, D. aurantiacus, D. bicolor, D. bigelovii, D. bolanderi, D. brandegeei, D. brevipes, D. calycinus, D. cascadensis, D. clevelandii, D. clivicola, D. compactus, D. congdonii, D. constrictus, D. cusickii, D. cusickioides, D. deschutesensis, D. douglasii, D. fremontii, D. grandiflorus, D. graniticola, D. johnstonii, D. kelloggii, D. layneae, D. leptaleus, D. linearis, D. longiflorus, D. mephiticus, D. mohavensis, D. nanus, D. ovatus, D. parryi, D. parviflorus, D. pictus, D. pulchellus, D. puniceus, D. pygmaeus, D. rattanii, D. rupicola, D. rutilus, D. thompsonii, D. torreyi, D. traskiae, D. tricolor, D. vandenbergensis, D. viscidus

Synonyms

Eunanus layneae, Mimulus brachiatus, M. layneae

Mimulus jepsonii, M. microcarpus, M. nanus var. jepsonii

Name authority

(Greene) G. L. Nesom: Phytoneuron 2012-39: 29. (2012)

(A. L. Grant) G. L. Nesom: Phytoneuron 2012-39: 29. (2012)

Web links

Local floras: CA, OR
Local Web sites: CalFlora, CalPhotos, Flora NW, PNW Herbaria
WildflowerSearch
iNaturalist (observations)
LBJ Wildflower Center
SEINet
Plants of the World Online
Encyclopedia of Life
Wikipedia
Google Image Search

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

Choose a data source: Flora of North America, OregonFlora

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Diplacus layneae

Diplacus jepsonii

Diplacus layneae

Diplacus jepsonii