Diplacus layneae |
Diplacus deschutesensis |
|
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Layne's monkeyflower |
Deschutes monkeyflower |
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Habit | Herbs, annual, herbage usually drying dark. | Herbs, annual. |
Stems | erect, 30–160(–300) mm, nodes 3–6, glandular-puberulent to glandular-pubescent, hairs 0.2–0.8 mm. |
erect to erect-ascending, 40–150 mm, distal internodes 1–2 mm, minutely glandular-puberulent. |
Leaves | usually cauline, relatively even-sized; petiole weakly delimited; blade elliptic to narrowly elliptic, oblanceolate, elliptic-oblanceolate, or elliptic-lanceolate, 8–27(–35) × 2–8 mm, margins entire, rarely toothed, plane, apex rounded to obtuse, surfaces: proximals often glabrate, distals glandular-puberulent or glandular-pubescent. |
usually cauline, relatively even-sized or gradually larger distally; petiole absent, proximal base short petiole-like; blade broadly ovate or obovate to elliptic-ovate or elliptic-oblanceolate, 10–15(–25) × 4–13 mm, margins entire, plane, apex acuminate or cuspidate, surfaces glabrous or sparsely glandular-puberulent. |
Pedicels | 2–4(–5) mm in fruit. |
1–1.5 mm in fruit. |
Flowers | 2 per node, or 1 or 2 per node on 1 plant, chasmogamous. |
usually from proximalmost to distal nodes, 2 per node, or 1 or 2 per node on 1 plant, chasmogamous. |
Styles | glandular-puberulent. |
puberulent, at least on distal 1/2. |
Corollas | pinkish or pale to dark magenta or red-purple, each lobe usually with a faint to dark medial line extending 1/2 or less to tip, throat floor yellowish near base, mostly white with red-purple dots near mouth, palate ridges white, tube-throat 10–15 mm, limb (8–)10–16 mm diam., not bilabiate. |
light pink to magenta or rose purple, usually with a darker narrow line extending from throat onto each lobe midvein, throat yellow, palate ridges yellow, tube-throat 8–12 mm, limb 10–16 mm diam., bilabiate. |
Calyces | symmetrically attached to pedicels, not inflated in fruit, (5–)6–8(–9) mm, glandular-puberulent to glandular-pubescent, tube strongly plicate, lobes triangular, subequal, apex acute, ribs broad, darkened, blackish, thickened, strongly raised, intercostal areas whitish, membranous. |
symmetrically attached to pedicels, not inflated in fruit, 7–8 mm, glandular-puberulent, lobes subequal, apex linear-acuminate, ribs green distally, intercostal areas whitish. |
Capsules | 6–10(–13) mm. |
7–9 mm. |
Anthers | included, ciliate. |
included, glabrous or sparsely hispidulous. |
Stigmas | included, lobes unequal, abaxial 1.5 times adaxial. |
exserted, lobes subequal, abaxial slightly longer. |
2n | = 16. |
= 16. |
Diplacus layneae |
Diplacus deschutesensis |
|
Phenology | Flowering May–Aug. | Flowering Jun–Aug. |
Habitat | Road banks, serpentine, granitic sand, red clay, lava beds and volcanic soils, openings in chaparral, shallow dry streambeds or stream banks, burned or otherwise disturbed open areas. | Sandy and ashy soils, pumice sand and gravel, red clay slopes, hillsides, roadsides, bare areas, sagebrush, sagebrush-juniper, juniper, yellow pine and lodgepole pine forests. |
Elevation | (100–)400–2400 m. ((300–)1300–7900 ft.) | 700–1500 m. (2300–4900 ft.) |
Distribution |
CA
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OR
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Discussion | D. M. Thompson (2005) observed that two forms of Diplacus layneae co-occur from the Yosemite National Park area southward; one of these is recognized here as D. graniticola. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Diplacus deschutesensis is endemic to Crook, Deschutes, Jefferson, Klamath, Lake, and Wheeler counties of central Oregon. D. M. Thompson (2005) regarded these plants as a zone of stabilized hybrids, intermediate between Mimulus cusickii and typical M. nanus, the range just outside and west of the wider range of typical M. cusickii. In an earlier study that included both of the latter species, W. L. Ezell (1971, and by annotation in 1987) identified the same set of plants simply as M. cusickii, not associating them at all with M. nanus. A. L. Grant (1924, and by annotation of MO collections) identified them variously as either M. cusickii or M. ovatus. Thompson did not say what features of intermediacy he observed in the putative hybrids, but he did note that they produced leaves with acuminate-cuspidate apices and that they would key to M. cusickii. Leaves of Diplacus deschutesensis are broad with abruptly and sharply acuminate apices like those of D. cusickii, and the corolla coloration also is similar. The flowers (calyx length, corolla tube-throat length, limb width) and capsules of D. deschutesensis are considerably smaller, and the distal leaves are smaller with glabrous surfaces. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 437. | FNA vol. 17, p. 440. |
Parent taxa | Phrymaceae > Diplacus | Phrymaceae > Diplacus |
Sibling taxa | ||
Synonyms | Eunanus layneae, Mimulus brachiatus, M. layneae | |
Name authority | (Greene) G. L. Nesom: Phytoneuron 2012-39: 29. (2012) | G. L. Nesom: Phytoneuron 2013-65: 8, fig. 5. (2013) |
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