FNA: Diplacus layneae vs. Diplacus cusickioides

	Diplacus layneae	Diplacus cusickioides
	Layne's monkeyflower	Cusick's monkeyflower, Nesom's monkey-flower
Habit	Herbs, annual, herbage usually drying dark.	Herbs, annual.
Stems	erect, 30–160(–300) mm, nodes 3–6, glandular-puberulent to glandular-pubescent, hairs 0.2–0.8 mm.	erect to erect-ascending, (10–)30–240(–350) mm, distal internodes 1–5 mm, densely glandular-villous.
Leaves	usually cauline, relatively even-sized; petiole weakly delimited; blade elliptic to narrowly elliptic, oblanceolate, elliptic-oblanceolate, or elliptic-lanceolate, 8–27(–35) × 2–8 mm, margins entire, rarely toothed, plane, apex rounded to obtuse, surfaces: proximals often glabrate, distals glandular-puberulent or glandular-pubescent.	basal and cauline or usually cauline, relatively even-sized or gradually larger distally; petiole absent, proximal base short petiole-like, 1–5 mm; blade ovate to broadly elliptic-ovate, (10–)15–25(–35) × 4–16 mm, margins entire, plane, apex abruptly acuminate, surfaces densely glandular-villous.
Pedicels	2–4(–5) mm in fruit.	1–1.5 mm in fruit.
Flowers	2 per node, or 1 or 2 per node on 1 plant, chasmogamous.	2 per node, or 1 or 2 per node on 1 plant, chasmogamous.
Styles	glandular-puberulent.	puberulent, at least on distal 1/2.
Corollas	pinkish or pale to dark magenta or red-purple, each lobe usually with a faint to dark medial line extending 1/2 or less to tip, throat floor yellowish near base, mostly white with red-purple dots near mouth, palate ridges white, tube-throat 10–15 mm, limb (8–)10–16 mm diam., not bilabiate.	magenta or rose purple, tube yellow, throat usually yellow, throat and distal tube red-spotted on floor, palate ridges yellow, tube-throat 13–16(–19) mm, limb 14–26 mm diam., bilabiate.
Calyces	symmetrically attached to pedicels, not inflated in fruit, (5–)6–8(–9) mm, glandular-puberulent to glandular-pubescent, tube strongly plicate, lobes triangular, subequal, apex acute, ribs broad, darkened, blackish, thickened, strongly raised, intercostal areas whitish, membranous.	symmetrically attached to pedicels, not inflated in fruit, 7–12 mm, glabrous or minutely stipitate-glandular, lobes unequal, apex linear-acuminate, sharp-pointed, ribs green distally, intercostal areas whitish.
Capsules	6–10(–13) mm.	10–17 mm.
Anthers	included, ciliate.	included, glabrous or sparsely hirsutulous.
Stigmas	included, lobes unequal, abaxial 1.5 times adaxial.	exserted, lobes subequal (herkogamous).
2n	= 16.

	Diplacus layneae	Diplacus cusickioides
Phenology	Flowering May–Aug.	Flowering May–Aug.
Habitat	Road banks, serpentine, granitic sand, red clay, lava beds and volcanic soils, openings in chaparral, shallow dry streambeds or stream banks, burned or otherwise disturbed open areas.	Lava formations, steep slopes, roadsides, volcanic gravels, scree, ash.
Elevation	(100–)400–2400 m. ((300–)1300–7900 ft.)	(400–)600–1500(–2000) m. ((1300–)2000–4900(–6600) ft.)
Distribution	CA [WildflowerSearch map]	CA; ID; OR; WA [WildflowerSearch map]
Discussion	D. M. Thompson (2005) observed that two forms of Diplacus layneae co-occur from the Yosemite National Park area southward; one of these is recognized here as D. graniticola. (Discussion copyrighted by Flora of North America; reprinted with permission.)	Diplacus cusickioides occurs in western Idaho, eastern Oregon, Klickitat County, Washington, and apparently in a disjunct population system in Modoc County, California, on the east side of the Warner Mountains. (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 17, p. 437.	FNA vol. 17, p. 439.
Parent taxa	Phrymaceae > Diplacus	Phrymaceae > Diplacus
Sibling taxa	D. angustatus, D. aridus, D. aurantiacus, D. bicolor, D. bigelovii, D. bolanderi, D. brandegeei, D. brevipes, D. calycinus, D. cascadensis, D. clevelandii, D. clivicola, D. compactus, D. congdonii, D. constrictus, D. cusickii, D. cusickioides, D. deschutesensis, D. douglasii, D. fremontii, D. grandiflorus, D. graniticola, D. jepsonii, D. johnstonii, D. kelloggii, D. leptaleus, D. linearis, D. longiflorus, D. mephiticus, D. mohavensis, D. nanus, D. ovatus, D. parryi, D. parviflorus, D. pictus, D. pulchellus, D. puniceus, D. pygmaeus, D. rattanii, D. rupicola, D. rutilus, D. thompsonii, D. torreyi, D. traskiae, D. tricolor, D. vandenbergensis, D. viscidus	D. angustatus, D. aridus, D. aurantiacus, D. bicolor, D. bigelovii, D. bolanderi, D. brandegeei, D. brevipes, D. calycinus, D. cascadensis, D. clevelandii, D. clivicola, D. compactus, D. congdonii, D. constrictus, D. cusickii, D. deschutesensis, D. douglasii, D. fremontii, D. grandiflorus, D. graniticola, D. jepsonii, D. johnstonii, D. kelloggii, D. layneae, D. leptaleus, D. linearis, D. longiflorus, D. mephiticus, D. mohavensis, D. nanus, D. ovatus, D. parryi, D. parviflorus, D. pictus, D. pulchellus, D. puniceus, D. pygmaeus, D. rattanii, D. rupicola, D. rutilus, D. thompsonii, D. torreyi, D. traskiae, D. tricolor, D. vandenbergensis, D. viscidus
Synonyms	Eunanus layneae, Mimulus brachiatus, M. layneae
Name authority	(Greene) G. L. Nesom: Phytoneuron 2012-39: 29. (2012)	G. L. Nesom: Phytoneuron 2013-65: 6, figs. 3, 4. (2013)
Web links	Local floras: CA Local Web sites: CalFlora, CalPhotos WildflowerSearch iNaturalist (observations) LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	Local floras: CA, OR, WA Local Web sites: CalFlora, CalPhotos, Flora NW, PNW Herbaria WildflowerSearch iNaturalist (observations) LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Diplacus cusickioides

Layne's monkeyflower

Cusick's monkeyflower, Nesom's monkey-flower

Habit

Herbs, annual, herbage usually drying dark.

Herbs, annual.

Stems

erect, 30–160(–300) mm, nodes 3–6, glandular-puberulent to glandular-pubescent, hairs 0.2–0.8 mm.

erect to erect-ascending, (10–)30–240(–350) mm, distal internodes 1–5 mm, densely glandular-villous.

Leaves

usually cauline, relatively even-sized;

petiole weakly delimited;

blade elliptic to narrowly elliptic, oblanceolate, elliptic-oblanceolate, or elliptic-lanceolate, 8–27(–35) × 2–8 mm, margins entire, rarely toothed, plane, apex rounded to obtuse, surfaces: proximals often glabrate, distals glandular-puberulent or glandular-pubescent.

basal and cauline or usually cauline, relatively even-sized or gradually larger distally;

petiole absent, proximal base short petiole-like, 1–5 mm;

blade ovate to broadly elliptic-ovate, (10–)15–25(–35) × 4–16 mm, margins entire, plane, apex abruptly acuminate, surfaces densely glandular-villous.

Pedicels

2–4(–5) mm in fruit.

1–1.5 mm in fruit.

Flowers

2 per node, or 1 or 2 per node on 1 plant, chasmogamous.

Styles

glandular-puberulent.

puberulent, at least on distal 1/2.

Corollas

pinkish or pale to dark magenta or red-purple, each lobe usually with a faint to dark medial line extending 1/2 or less to tip, throat floor yellowish near base, mostly white with red-purple dots near mouth, palate ridges white, tube-throat 10–15 mm, limb (8–)10–16 mm diam., not bilabiate.

magenta or rose purple, tube yellow, throat usually yellow, throat and distal tube red-spotted on floor, palate ridges yellow, tube-throat 13–16(–19) mm, limb 14–26 mm diam., bilabiate.

Calyces

symmetrically attached to pedicels, not inflated in fruit, (5–)6–8(–9) mm, glandular-puberulent to glandular-pubescent, tube strongly plicate, lobes triangular, subequal, apex acute, ribs broad, darkened, blackish, thickened, strongly raised, intercostal areas whitish, membranous.

symmetrically attached to pedicels, not inflated in fruit, 7–12 mm, glabrous or minutely stipitate-glandular, lobes unequal, apex linear-acuminate, sharp-pointed, ribs green distally, intercostal areas whitish.

Capsules

6–10(–13) mm.

10–17 mm.

Anthers

included, ciliate.

included, glabrous or sparsely hirsutulous.

Stigmas

included, lobes unequal, abaxial 1.5 times adaxial.

exserted, lobes subequal (herkogamous).

= 16.

Diplacus layneae

Diplacus cusickioides

Phenology

Flowering May–Aug.

Habitat

Road banks, serpentine, granitic sand, red clay, lava beds and volcanic soils, openings in chaparral, shallow dry streambeds or stream banks, burned or otherwise disturbed open areas.

Lava formations, steep slopes, roadsides, volcanic gravels, scree, ash.

Elevation

(100–)400–2400 m. ((300–)1300–7900 ft.)

(400–)600–1500(–2000) m. ((1300–)2000–4900(–6600) ft.)

Distribution

[WildflowerSearch map]

CA; ID; OR; WA

[WildflowerSearch map]

Discussion

D. M. Thompson (2005) observed that two forms of Diplacus layneae co-occur from the Yosemite National Park area southward; one of these is recognized here as D. graniticola.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Diplacus cusickioides occurs in western Idaho, eastern Oregon, Klickitat County, Washington, and apparently in a disjunct population system in Modoc County, California, on the east side of the Warner Mountains.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 17, p. 437.

FNA vol. 17, p. 439.

Parent taxa

Phrymaceae > Diplacus

Sibling taxa

D. angustatus, D. aridus, D. aurantiacus, D. bicolor, D. bigelovii, D. bolanderi, D. brandegeei, D. brevipes, D. calycinus, D. cascadensis, D. clevelandii, D. clivicola, D. compactus, D. congdonii, D. constrictus, D. cusickii, D. cusickioides, D. deschutesensis, D. douglasii, D. fremontii, D. grandiflorus, D. graniticola, D. jepsonii, D. johnstonii, D. kelloggii, D. leptaleus, D. linearis, D. longiflorus, D. mephiticus, D. mohavensis, D. nanus, D. ovatus, D. parryi, D. parviflorus, D. pictus, D. pulchellus, D. puniceus, D. pygmaeus, D. rattanii, D. rupicola, D. rutilus, D. thompsonii, D. torreyi, D. traskiae, D. tricolor, D. vandenbergensis, D. viscidus

D. angustatus, D. aridus, D. aurantiacus, D. bicolor, D. bigelovii, D. bolanderi, D. brandegeei, D. brevipes, D. calycinus, D. cascadensis, D. clevelandii, D. clivicola, D. compactus, D. congdonii, D. constrictus, D. cusickii, D. deschutesensis, D. douglasii, D. fremontii, D. grandiflorus, D. graniticola, D. jepsonii, D. johnstonii, D. kelloggii, D. layneae, D. leptaleus, D. linearis, D. longiflorus, D. mephiticus, D. mohavensis, D. nanus, D. ovatus, D. parryi, D. parviflorus, D. pictus, D. pulchellus, D. puniceus, D. pygmaeus, D. rattanii, D. rupicola, D. rutilus, D. thompsonii, D. torreyi, D. traskiae, D. tricolor, D. vandenbergensis, D. viscidus

Synonyms

Eunanus layneae, Mimulus brachiatus, M. layneae

Name authority

(Greene) G. L. Nesom: Phytoneuron 2012-39: 29. (2012)

G. L. Nesom: Phytoneuron 2013-65: 6, figs. 3, 4. (2013)

Web links

Local floras: CA, OR, WA
Local Web sites: CalFlora, CalPhotos, Flora NW, PNW Herbaria
WildflowerSearch
iNaturalist (observations)
LBJ Wildflower Center
SEINet
Plants of the World Online
Encyclopedia of Life
Wikipedia
Google Image Search

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

Choose a data source: Flora of North America, OregonFlora, UW Burke Herbarium

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Diplacus layneae

Diplacus cusickioides

Diplacus layneae

Diplacus cusickioides