Diplacus layneae |
Diplacus cusickii |
|
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Layne's monkeyflower |
Cusick's monkey-flower |
|
Habit | Herbs, annual, herbage usually drying dark. | Herbs, annual. |
Stems | erect, 30–160(–300) mm, nodes 3–6, glandular-puberulent to glandular-pubescent, hairs 0.2–0.8 mm. |
erect to erect-ascending, 10–80 mm, distal internodes 2–20 mm, minutely glandular-puberulent. |
Leaves | usually cauline, relatively even-sized; petiole weakly delimited; blade elliptic to narrowly elliptic, oblanceolate, elliptic-oblanceolate, or elliptic-lanceolate, 8–27(–35) × 2–8 mm, margins entire, rarely toothed, plane, apex rounded to obtuse, surfaces: proximals often glabrate, distals glandular-puberulent or glandular-pubescent. |
basal and cauline or usually cauline, relatively even-sized or gradually larger distally; petiole absent, proximal base short petiole-like, 1–5 mm; blade ovate to broadly elliptic-ovate, (10–)15–25(–35) × 4–17 mm, margins entire, plane, apex acuminate, surfaces glabrous or adaxial minutely glandular-puberulent. |
Pedicels | 2–4(–5) mm in fruit. |
1–1.5 mm in fruit. |
Flowers | 2 per node, or 1 or 2 per node on 1 plant, chasmogamous. |
2 per node, or 1 or 2 per node on 1 plant, chasmogamous. |
Styles | glandular-puberulent. |
pubescent, at least on distal 1/2. |
Corollas | pinkish or pale to dark magenta or red-purple, each lobe usually with a faint to dark medial line extending 1/2 or less to tip, throat floor yellowish near base, mostly white with red-purple dots near mouth, palate ridges white, tube-throat 10–15 mm, limb (8–)10–16 mm diam., not bilabiate. |
magenta or rose purple, tube yellow, throat yellow, throat and distal tube red-spotted on floor, palate ridges yellow, tube-throat 13–16(–19) mm, limb 16–24 mm diam., bilabiate. |
Calyces | symmetrically attached to pedicels, not inflated in fruit, (5–)6–8(–9) mm, glandular-puberulent to glandular-pubescent, tube strongly plicate, lobes triangular, subequal, apex acute, ribs broad, darkened, blackish, thickened, strongly raised, intercostal areas whitish, membranous. |
symmetrically attached to pedicels, not inflated in fruit, 7–10 mm, glabrous or minutely stipitate-glandular, lobes unequal, apex linear-acuminate, sharp-pointed, ribs green distally, intercostal areas whitish. |
Capsules | 6–10(–13) mm. |
10–15 mm. |
Anthers | included, ciliate. |
included, glabrous or sparsely hirsutulous. |
Stigmas | included, lobes unequal, abaxial 1.5 times adaxial. |
exserted, lobes subequal. |
2n | = 16. |
|
Diplacus layneae |
Diplacus cusickii |
|
Phenology | Flowering May–Aug. | Flowering May–Jul. |
Habitat | Road banks, serpentine, granitic sand, red clay, lava beds and volcanic soils, openings in chaparral, shallow dry streambeds or stream banks, burned or otherwise disturbed open areas. | Slopes, canyons, washes, ditches, sand talus, diatomaceous slopes, basalt outcrops, black volcanic gravel, volcanic ash and sand, sagebrush areas. |
Elevation | (100–)400–2400 m. ((300–)1300–7900 ft.) | 800–1000 m. (2600–3300 ft.) |
Distribution |
CA
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ID; OR
|
Discussion | D. M. Thompson (2005) observed that two forms of Diplacus layneae co-occur from the Yosemite National Park area southward; one of these is recognized here as D. graniticola. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Diplacus cusickii is endemic to northern Malheur County, Oregon, and along the Snake River in Ada and Owyhee counties, Idaho. Its narrow geographic range reflects the segregation of the more widely distributed D. cusickioides. D. M. Thompson (2005) noted that collections in northern Malheur County, Oregon, were intermediate between Mimulus cusickii and M. nanus; the type of M. cusickii is from this area and is among the narrowly endemic, supposedly putative intermediates (G. L. Nesom 2013c). These plants have abruptly and sharply acuminate leaf apices like the more widespread form traditionally identified as Diplacus cusickii; they differ in having glabrous leaf surfaces. Typical D. nanus occurs in close sympatry, without intergradation, with the populations in northern Malheur County. Because of their distinctive morphology and coherent geography, the northern Malheur County plants are reasonably recognized as a distinct species. The more widely distributed form formerly identified as D. cusickii now is identified as D. cusickioides. Populations of D. cusickii in the narrow sense along the Snake River in Ada and Owyhee counties, Idaho, may have dispersed there from the Oregon center. Some plants of D. cusickioides in the Leslie Gulch area of east-central Malheur County have somewhat reduced vestiture, approaching that of D. cusickii. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 437. | FNA vol. 17, p. 439. |
Parent taxa | Phrymaceae > Diplacus | Phrymaceae > Diplacus |
Sibling taxa | ||
Synonyms | Eunanus layneae, Mimulus brachiatus, M. layneae | Eunanus cusickii, Mimulus cusickii |
Name authority | (Greene) G. L. Nesom: Phytoneuron 2012-39: 29. (2012) | (Greene) G. L. Nesom: Phytoneuron 2012-39: 28. (2012) |
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