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Layne's monkeyflower

Congdon's monkeyflower

Habit Herbs, annual, herbage usually drying dark. Herbs, annual.
Stems

erect, 30–160(–300) mm, nodes 3–6, glandular-puberulent to glandular-pubescent, hairs 0.2–0.8 mm.

erect to ascending, 0–100(–120) mm, glandular-puberulent and/or glandular-pubescent.

Leaves

usually cauline, relatively even-sized;

petiole weakly delimited;

blade elliptic to narrowly elliptic, oblanceolate, elliptic-oblanceolate, or elliptic-lanceolate, 8–27(–35) × 2–8 mm, margins entire, rarely toothed, plane, apex rounded to obtuse, surfaces: proximals often glabrate, distals glandular-puberulent or glandular-pubescent.

usually cauline, relatively even-sized;

petiole absent, larger with petiole-like extension;

blade oblanceolate to elliptic, (4–)8–32(–37) × (1.5–)2.5–14(–18) mm, margins entire or crenate, plane, apex obtuse, surfaces glabrous or puberulent and/or pilose adaxially.

Pedicels

2–4(–5) mm in fruit.

(1–)2–5 mm in fruit, usually twisting to invert calyx.

Flowers

2 per node, or 1 or 2 per node on 1 plant, chasmogamous.

2 per node, or 1 or 2 per node on 1 plant, chasmogamous or cleistogamous.

Styles

glandular-puberulent.

glandular-pubescent.

Corollas

pinkish or pale to dark magenta or red-purple, each lobe usually with a faint to dark medial line extending 1/2 or less to tip, throat floor yellowish near base, mostly white with red-purple dots near mouth, palate ridges white, tube-throat 10–15 mm, limb (8–)10–16 mm diam., not bilabiate.

throat whitish to dark magenta, not golden yellow at base, lobes magenta, without markings or with dark magenta dots and, sometimes, some yellow at bases of abaxial lobes, palate ridges absent or purple, tube-throat 8–30 mm, limb 3–9 mm diam., bilabiate, abaxial lobe usually smaller than adaxial.

Calyces

symmetrically attached to pedicels, not inflated in fruit, (5–)6–8(–9) mm, glandular-puberulent to glandular-pubescent, tube strongly plicate, lobes triangular, subequal, apex acute, ribs broad, darkened, blackish, thickened, strongly raised, intercostal areas whitish, membranous.

distinctly asymmetrically attached to pedicel, inflated in fruit, 5–14 mm, sparsely pilose, lobes subequal, apex obtuse, ribs green to purplish, intercostal areas whitish.

Capsules

6–10(–13) mm.

4–8.5 mm, indehiscent.

Anthers

included, ciliate.

(distal pair) exserted, glabrous.

Stigmas

included, lobes unequal, abaxial 1.5 times adaxial.

exserted, lobes unequal, abaxial 3–4 times adaxial.

2n

= 16.

= 18.

Diplacus layneae

Diplacus congdonii

Phenology Flowering May–Aug. Flowering Feb–May.
Habitat Road banks, serpentine, granitic sand, red clay, lava beds and volcanic soils, openings in chaparral, shallow dry streambeds or stream banks, burned or otherwise disturbed open areas. Serpentine soils, periphery of granite outcrops, disturbed hillsides, soil from decomposed granite, near water runoff areas, away from seeps or other areas with prolonged surface moisture.
Elevation (100–)400–2400 m. ((300–)1300–7900 ft.) 100–1100(–1700) m. (300–3600(–5600) ft.)
Distribution
from FNA
CA
[WildflowerSearch map]
from FNA
CA
[WildflowerSearch map]
Discussion

D. M. Thompson (2005) observed that two forms of Diplacus layneae co-occur from the Yosemite National Park area southward; one of these is recognized here as D. graniticola.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Diplacus congdonii occurs in the Sierra Nevada Foothills and mountain ranges along the coast.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 437. FNA vol. 17, p. 445.
Parent taxa Phrymaceae > Diplacus Phrymaceae > Diplacus
Sibling taxa
D. angustatus, D. aridus, D. aurantiacus, D. bicolor, D. bigelovii, D. bolanderi, D. brandegeei, D. brevipes, D. calycinus, D. cascadensis, D. clevelandii, D. clivicola, D. compactus, D. congdonii, D. constrictus, D. cusickii, D. cusickioides, D. deschutesensis, D. douglasii, D. fremontii, D. grandiflorus, D. graniticola, D. jepsonii, D. johnstonii, D. kelloggii, D. leptaleus, D. linearis, D. longiflorus, D. mephiticus, D. mohavensis, D. nanus, D. ovatus, D. parryi, D. parviflorus, D. pictus, D. pulchellus, D. puniceus, D. pygmaeus, D. rattanii, D. rupicola, D. rutilus, D. thompsonii, D. torreyi, D. traskiae, D. tricolor, D. vandenbergensis, D. viscidus
D. angustatus, D. aridus, D. aurantiacus, D. bicolor, D. bigelovii, D. bolanderi, D. brandegeei, D. brevipes, D. calycinus, D. cascadensis, D. clevelandii, D. clivicola, D. compactus, D. constrictus, D. cusickii, D. cusickioides, D. deschutesensis, D. douglasii, D. fremontii, D. grandiflorus, D. graniticola, D. jepsonii, D. johnstonii, D. kelloggii, D. layneae, D. leptaleus, D. linearis, D. longiflorus, D. mephiticus, D. mohavensis, D. nanus, D. ovatus, D. parryi, D. parviflorus, D. pictus, D. pulchellus, D. puniceus, D. pygmaeus, D. rattanii, D. rupicola, D. rutilus, D. thompsonii, D. torreyi, D. traskiae, D. tricolor, D. vandenbergensis, D. viscidus
Synonyms Eunanus layneae, Mimulus brachiatus, M. layneae Mimulus congdonii
Name authority (Greene) G. L. Nesom: Phytoneuron 2012-39: 29. (2012) (B. L. Robinson) G. L. Nesom: Phytoneuron 2012-39: 32. (2012)
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