FNA: Diplacus layneae vs. Diplacus cascadensis

	Diplacus layneae	Diplacus cascadensis
	Layne's monkeyflower	Cascade monkeyflower, Cascades monkeyflower
Habit	Herbs, annual, herbage usually drying dark.	Herbs, annual.
Stems	erect, 30–160(–300) mm, nodes 3–6, glandular-puberulent to glandular-pubescent, hairs 0.2–0.8 mm.	erect to ascending-erect, 20–100 mm, distal internodes 1–4 mm, short glandular-villous to glandular-puberulent.
Leaves	usually cauline, relatively even-sized; petiole weakly delimited; blade elliptic to narrowly elliptic, oblanceolate, elliptic-oblanceolate, or elliptic-lanceolate, 8–27(–35) × 2–8 mm, margins entire, rarely toothed, plane, apex rounded to obtuse, surfaces: proximals often glabrate, distals glandular-puberulent or glandular-pubescent.	usually cauline, relatively even-sized; petiole present proximally, absent distally; blade elliptic-spatulate to obovate or broadly oblanceolate, 10–22 × 2–10 mm, margins entire, plane, apex obtuse to rounded-acute, surfaces minutely glandular-puberulent.
Pedicels	2–4(–5) mm in fruit.	1–3 mm in fruit.
Flowers	2 per node, or 1 or 2 per node on 1 plant, chasmogamous.	2 per node, or 1 or 2 per node on 1 plant, chasmogamous.
Styles	glandular-puberulent.	apparently glabrous.
Corollas	pinkish or pale to dark magenta or red-purple, each lobe usually with a faint to dark medial line extending 1/2 or less to tip, throat floor yellowish near base, mostly white with red-purple dots near mouth, palate ridges white, tube-throat 10–15 mm, limb (8–)10–16 mm diam., not bilabiate.	magenta to purplish, usually with a darker narrow line extending from throat onto each lobe midvein, palate ridges yellow with red spots, throat floor glabrous, tube 1.1–1.9 mm diam. at filament insertion, tube-throat 8–10 mm, limb 7–11 mm diam., bilabiate.
Calyces	symmetrically attached to pedicels, not inflated in fruit, (5–)6–8(–9) mm, glandular-puberulent to glandular-pubescent, tube strongly plicate, lobes triangular, subequal, apex acute, ribs broad, darkened, blackish, thickened, strongly raised, intercostal areas whitish, membranous.	symmetrically attached to pedicels, not inflated in fruit, (4–)5–7 mm, minutely glandular-puberulent, lobes subequal, apex acute to acuminate, ribs dark green or reddish, intercostal areas whitish.
Capsules	6–10(–13) mm.	5–8(–9) mm.
Anthers	included, ciliate.	(distal pair) exserted, minutely viscid-villosulous.
Stigmas	included, lobes unequal, abaxial 1.5 times adaxial.	exserted, lobes usually subequal.
2n	= 16.	= 16.

	Diplacus layneae	Diplacus cascadensis
Phenology	Flowering May–Aug.	Flowering Jun–Aug.
Habitat	Road banks, serpentine, granitic sand, red clay, lava beds and volcanic soils, openings in chaparral, shallow dry streambeds or stream banks, burned or otherwise disturbed open areas.	Open pumice flats, scree slopes, sandy soils, juniper-sagebrush, juniper, pine-juniper, yellow pine, lodgepole pine forests.
Elevation	(100–)400–2400 m. ((300–)1300–7900 ft.)	1400–2400(–2600) m. (4600–7900(–8500) ft.)
Distribution	CA [WildflowerSearch map]	OR
Discussion	D. M. Thompson (2005) observed that two forms of Diplacus layneae co-occur from the Yosemite National Park area southward; one of these is recognized here as D. graniticola. (Discussion copyrighted by Flora of North America; reprinted with permission.)	Diplacus cascadensis is known from Deschutes, Klamath, and Lake counties. Plants of this species have been identified as D. nanus (similar in its purplish leaves congested on crowded distal nodes, minutely glandular-puberulent vestiture, and purplish and strongly bilabiate corollas), but they differ from D. nanus in their broader distal leaves, shorter calyces, shorter corollas with glabrous throats and magenta tubes, and shorter capsules. (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 17, p. 437.	FNA vol. 17, p. 441.
Parent taxa	Phrymaceae > Diplacus	Phrymaceae > Diplacus
Sibling taxa	D. angustatus, D. aridus, D. aurantiacus, D. bicolor, D. bigelovii, D. bolanderi, D. brandegeei, D. brevipes, D. calycinus, D. cascadensis, D. clevelandii, D. clivicola, D. compactus, D. congdonii, D. constrictus, D. cusickii, D. cusickioides, D. deschutesensis, D. douglasii, D. fremontii, D. grandiflorus, D. graniticola, D. jepsonii, D. johnstonii, D. kelloggii, D. leptaleus, D. linearis, D. longiflorus, D. mephiticus, D. mohavensis, D. nanus, D. ovatus, D. parryi, D. parviflorus, D. pictus, D. pulchellus, D. puniceus, D. pygmaeus, D. rattanii, D. rupicola, D. rutilus, D. thompsonii, D. torreyi, D. traskiae, D. tricolor, D. vandenbergensis, D. viscidus	D. angustatus, D. aridus, D. aurantiacus, D. bicolor, D. bigelovii, D. bolanderi, D. brandegeei, D. brevipes, D. calycinus, D. clevelandii, D. clivicola, D. compactus, D. congdonii, D. constrictus, D. cusickii, D. cusickioides, D. deschutesensis, D. douglasii, D. fremontii, D. grandiflorus, D. graniticola, D. jepsonii, D. johnstonii, D. kelloggii, D. layneae, D. leptaleus, D. linearis, D. longiflorus, D. mephiticus, D. mohavensis, D. nanus, D. ovatus, D. parryi, D. parviflorus, D. pictus, D. pulchellus, D. puniceus, D. pygmaeus, D. rattanii, D. rupicola, D. rutilus, D. thompsonii, D. torreyi, D. traskiae, D. tricolor, D. vandenbergensis, D. viscidus
Synonyms	Eunanus layneae, Mimulus brachiatus, M. layneae
Name authority	(Greene) G. L. Nesom: Phytoneuron 2012-39: 29. (2012)	G. L. Nesom: Phytoneuron 2013-65: 13, figs. 8, 9. (2013)
Web links	Local floras: CA Local Web sites: CalFlora, CalPhotos WildflowerSearch iNaturalist (observations) LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	Local floras: OR Local Web sites: Flora NW, PNW Herbaria WildflowerSearch iNaturalist (observations) LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Diplacus cascadensis

Layne's monkeyflower

Cascade monkeyflower, Cascades monkeyflower

Habit

Herbs, annual, herbage usually drying dark.

Herbs, annual.

Stems

erect, 30–160(–300) mm, nodes 3–6, glandular-puberulent to glandular-pubescent, hairs 0.2–0.8 mm.

erect to ascending-erect, 20–100 mm, distal internodes 1–4 mm, short glandular-villous to glandular-puberulent.

Leaves

usually cauline, relatively even-sized;

petiole weakly delimited;

blade elliptic to narrowly elliptic, oblanceolate, elliptic-oblanceolate, or elliptic-lanceolate, 8–27(–35) × 2–8 mm, margins entire, rarely toothed, plane, apex rounded to obtuse, surfaces: proximals often glabrate, distals glandular-puberulent or glandular-pubescent.

usually cauline, relatively even-sized;

petiole present proximally, absent distally;

blade elliptic-spatulate to obovate or broadly oblanceolate, 10–22 × 2–10 mm, margins entire, plane, apex obtuse to rounded-acute, surfaces minutely glandular-puberulent.

Pedicels

2–4(–5) mm in fruit.

1–3 mm in fruit.

Flowers

2 per node, or 1 or 2 per node on 1 plant, chasmogamous.

Styles

glandular-puberulent.

apparently glabrous.

Corollas

pinkish or pale to dark magenta or red-purple, each lobe usually with a faint to dark medial line extending 1/2 or less to tip, throat floor yellowish near base, mostly white with red-purple dots near mouth, palate ridges white, tube-throat 10–15 mm, limb (8–)10–16 mm diam., not bilabiate.

magenta to purplish, usually with a darker narrow line extending from throat onto each lobe midvein, palate ridges yellow with red spots, throat floor glabrous, tube 1.1–1.9 mm diam. at filament insertion, tube-throat 8–10 mm, limb 7–11 mm diam., bilabiate.

Calyces

symmetrically attached to pedicels, not inflated in fruit, (5–)6–8(–9) mm, glandular-puberulent to glandular-pubescent, tube strongly plicate, lobes triangular, subequal, apex acute, ribs broad, darkened, blackish, thickened, strongly raised, intercostal areas whitish, membranous.

symmetrically attached to pedicels, not inflated in fruit, (4–)5–7 mm, minutely glandular-puberulent, lobes subequal, apex acute to acuminate, ribs dark green or reddish, intercostal areas whitish.

Capsules

6–10(–13) mm.

5–8(–9) mm.

Anthers

included, ciliate.

(distal pair) exserted, minutely viscid-villosulous.

Stigmas

included, lobes unequal, abaxial 1.5 times adaxial.

exserted, lobes usually subequal.

= 16.

Diplacus layneae

Diplacus cascadensis

Phenology

Flowering May–Aug.

Flowering Jun–Aug.

Habitat

Road banks, serpentine, granitic sand, red clay, lava beds and volcanic soils, openings in chaparral, shallow dry streambeds or stream banks, burned or otherwise disturbed open areas.

Open pumice flats, scree slopes, sandy soils, juniper-sagebrush, juniper, pine-juniper, yellow pine, lodgepole pine forests.

Elevation

(100–)400–2400 m. ((300–)1300–7900 ft.)

1400–2400(–2600) m. (4600–7900(–8500) ft.)

Distribution

[WildflowerSearch map]

Discussion

D. M. Thompson (2005) observed that two forms of Diplacus layneae co-occur from the Yosemite National Park area southward; one of these is recognized here as D. graniticola.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Diplacus cascadensis is known from Deschutes, Klamath, and Lake counties. Plants of this species have been identified as D. nanus (similar in its purplish leaves congested on crowded distal nodes, minutely glandular-puberulent vestiture, and purplish and strongly bilabiate corollas), but they differ from D. nanus in their broader distal leaves, shorter calyces, shorter corollas with glabrous throats and magenta tubes, and shorter capsules.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 17, p. 437.

FNA vol. 17, p. 441.

Parent taxa

Phrymaceae > Diplacus

Sibling taxa

D. angustatus, D. aridus, D. aurantiacus, D. bicolor, D. bigelovii, D. bolanderi, D. brandegeei, D. brevipes, D. calycinus, D. cascadensis, D. clevelandii, D. clivicola, D. compactus, D. congdonii, D. constrictus, D. cusickii, D. cusickioides, D. deschutesensis, D. douglasii, D. fremontii, D. grandiflorus, D. graniticola, D. jepsonii, D. johnstonii, D. kelloggii, D. leptaleus, D. linearis, D. longiflorus, D. mephiticus, D. mohavensis, D. nanus, D. ovatus, D. parryi, D. parviflorus, D. pictus, D. pulchellus, D. puniceus, D. pygmaeus, D. rattanii, D. rupicola, D. rutilus, D. thompsonii, D. torreyi, D. traskiae, D. tricolor, D. vandenbergensis, D. viscidus

D. angustatus, D. aridus, D. aurantiacus, D. bicolor, D. bigelovii, D. bolanderi, D. brandegeei, D. brevipes, D. calycinus, D. clevelandii, D. clivicola, D. compactus, D. congdonii, D. constrictus, D. cusickii, D. cusickioides, D. deschutesensis, D. douglasii, D. fremontii, D. grandiflorus, D. graniticola, D. jepsonii, D. johnstonii, D. kelloggii, D. layneae, D. leptaleus, D. linearis, D. longiflorus, D. mephiticus, D. mohavensis, D. nanus, D. ovatus, D. parryi, D. parviflorus, D. pictus, D. pulchellus, D. puniceus, D. pygmaeus, D. rattanii, D. rupicola, D. rutilus, D. thompsonii, D. torreyi, D. traskiae, D. tricolor, D. vandenbergensis, D. viscidus

Synonyms

Eunanus layneae, Mimulus brachiatus, M. layneae

Name authority

(Greene) G. L. Nesom: Phytoneuron 2012-39: 29. (2012)

G. L. Nesom: Phytoneuron 2013-65: 13, figs. 8, 9. (2013)

Web links

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

Choose a data source: Flora of North America, OregonFlora

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Flora of North America species comparison

Diplacus layneae

Diplacus cascadensis

Diplacus layneae

Diplacus cascadensis