Diplacus layneae |
Diplacus calycinus |
|
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Layne's monkeyflower |
Kaweah River bush monkeyflower, rock bush monkeyflower |
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Habit | Herbs, annual, herbage usually drying dark. | Subshrubs. |
Stems | erect, 30–160(–300) mm, nodes 3–6, glandular-puberulent to glandular-pubescent, hairs 0.2–0.8 mm. |
erect, 150–1500 mm, glandular-pubescent to viscid-villous. |
Leaves | usually cauline, relatively even-sized; petiole weakly delimited; blade elliptic to narrowly elliptic, oblanceolate, elliptic-oblanceolate, or elliptic-lanceolate, 8–27(–35) × 2–8 mm, margins entire, rarely toothed, plane, apex rounded to obtuse, surfaces: proximals often glabrate, distals glandular-puberulent or glandular-pubescent. |
cauline, relatively even-sized; petiole absent; blade elliptic-lanceolate or lanceolate to narrowly lanceolate or linear-lanceolate, sometimes narrowly oblong, 20–75(–100) × 4–20(–28) mm, margins entire or shallowly crenate, plane or revolute, apex acute to obtuse, abaxial surfaces moderately villous, hairs unbranched, vitreous, adaxial glabrous. |
Pedicels | 2–4(–5) mm in fruit. |
3–5 mm in fruit. |
Flowers | 2 per node, or 1 or 2 per node on 1 plant, chasmogamous. |
2 per node, chasmogamous. |
Styles | glandular-puberulent. |
minutely glandular. |
Corollas | pinkish or pale to dark magenta or red-purple, each lobe usually with a faint to dark medial line extending 1/2 or less to tip, throat floor yellowish near base, mostly white with red-purple dots near mouth, palate ridges white, tube-throat 10–15 mm, limb (8–)10–16 mm diam., not bilabiate. |
usually pale yellow or cream to yellow, not spotted or striped, palate ridges yellow to golden yellow, tube-throat 35–42 mm, limb 20–30 mm diam., bilabiate, lobes oblong, apex of adaxial 2 each shallowly, asymmetrically incised. |
Calyces | symmetrically attached to pedicels, not inflated in fruit, (5–)6–8(–9) mm, glandular-puberulent to glandular-pubescent, tube strongly plicate, lobes triangular, subequal, apex acute, ribs broad, darkened, blackish, thickened, strongly raised, intercostal areas whitish, membranous. |
not inflated in fruit, 28–40 mm, densely glandular-pubescent to short glandular-villous, tube slightly dilated distally, lobes unequal, apex acute, ribs green, intercostal areas light green. |
Capsules | 6–10(–13) mm. |
25–35 mm. |
Anthers | included, ciliate. |
exserted, glabrous. |
Stigmas | included, lobes unequal, abaxial 1.5 times adaxial. |
exserted, lobes equal. |
2n | = 16. |
= 20. |
Diplacus layneae |
Diplacus calycinus |
|
Phenology | Flowering May–Aug. | Flowering Apr–Jul. |
Habitat | Road banks, serpentine, granitic sand, red clay, lava beds and volcanic soils, openings in chaparral, shallow dry streambeds or stream banks, burned or otherwise disturbed open areas. | Granite outcrops, boulders, rocky gullies. |
Elevation | (100–)400–2400 m. ((300–)1300–7900 ft.) | (300–)700–2200 m. ((1000–)2300–7200 ft.) |
Distribution |
CA
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CA
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Discussion | D. M. Thompson (2005) observed that two forms of Diplacus layneae co-occur from the Yosemite National Park area southward; one of these is recognized here as D. graniticola. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Although first described as a separate species, Diplacus calycinus has more recently been treated at subspecific or varietal rank (A. L. Grant 1924; F. W. Pennell 1951; P. A. Munz and D. D. Keck 1973). D. M. Thompson (2005) included both D. calycinus and D. longiflorus within his concept of Mimulus aurantiacus var. pubescens (Torrey) D. M. Thompson. He did not reference the study of sect. Diplacus by M. C. Tulig (2000), but results from the Tulig morphometric analyses indicated that D. calycinus is distinct from D. longiflorus, especially in corolla length, corolla tube length, and style length. The type of Diplacus calycinus is from Tulare County, and the concept of the species is perhaps best restricted to the Sierran population system in Fresno, Kern, and Tulare counties, disjunct from D. longiflorus, which occurs primarily in coastal counties. The Sierran system is characterized by distinct abaxial leaf vestiture; the hairs are unbranched, broad, and vitreous, compared to the branched, thinner, and dull hairs of D. longiflorus. Plants of D. calycinus parapatric with D. longiflorus also show a tendency toward the characteristic vestiture and also have lighter-colored (but more variable in color) corollas with narrower but slightly shorter tubes. Intergradation between D. calycinus and D. longiflorus occurs in the region connecting the San Bernardino and San Gabriel mountains in San Bernardino County. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 437. | FNA vol. 17, p. 449. |
Parent taxa | Phrymaceae > Diplacus | Phrymaceae > Diplacus |
Sibling taxa | ||
Synonyms | Eunanus layneae, Mimulus brachiatus, M. layneae | D. longiflorus var. calycinus, Mimulus longiflorus subsp. calycinus, M. longiflorus var. calycinus |
Name authority | (Greene) G. L. Nesom: Phytoneuron 2012-39: 29. (2012) | Eastwood: Bot. Gaz. 41: 287. (1906) |
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