FNA: Diplacus layneae vs. Diplacus brandegeei

	Diplacus layneae	Diplacus brandegeei
	Layne's monkeyflower	Santa Cruz Island monkeyflower, Santa Cruz Island or broadleaf monkeyflower
Habit	Herbs, annual, herbage usually drying dark.	Herbs, annual.
Stems	erect, 30–160(–300) mm, nodes 3–6, glandular-puberulent to glandular-pubescent, hairs 0.2–0.8 mm.	erect, 10–100 mm, glandular-puberulent to glandular-pubescent.
Leaves	usually cauline, relatively even-sized; petiole weakly delimited; blade elliptic to narrowly elliptic, oblanceolate, elliptic-oblanceolate, or elliptic-lanceolate, 8–27(–35) × 2–8 mm, margins entire, rarely toothed, plane, apex rounded to obtuse, surfaces: proximals often glabrate, distals glandular-puberulent or glandular-pubescent.	mostly cauline, relatively even-sized; petiole absent, larger with petiole-like extension; blade narrowly to broadly ovate to broadly elliptic, (4–)8–39 × (2–)4–23 mm, margins entire or crenate, plane, proximal 1/2 usually ciliate, apex obtuse, surfaces glandular-pubescent.
Pedicels	2–4(–5) mm in fruit.	1.5–4 mm in fruit, not twisting to invert calyx.
Flowers	2 per node, or 1 or 2 per node on 1 plant, chasmogamous.	2 per node, or 1 or 2 per node on 1 plant, chasmogamous.
Styles	glandular-puberulent.	pubescent distally.
Corollas	pinkish or pale to dark magenta or red-purple, each lobe usually with a faint to dark medial line extending 1/2 or less to tip, throat floor yellowish near base, mostly white with red-purple dots near mouth, palate ridges white, tube-throat 10–15 mm, limb (8–)10–16 mm diam., not bilabiate.	throat magenta, golden yellow at base, limb magenta, palate ridges golden yellow, purple-speckled, tube-throat 12–19 mm, limb 5–11 mm diam., bilabiate, abaxial lip smaller than adaxial.
Calyces	symmetrically attached to pedicels, not inflated in fruit, (5–)6–8(–9) mm, glandular-puberulent to glandular-pubescent, tube strongly plicate, lobes triangular, subequal, apex acute, ribs broad, darkened, blackish, thickened, strongly raised, intercostal areas whitish, membranous.	distinctly asymmetrically attached to pedicel, inflated in fruit, 8–13 mm, glandular-pubescent, lobes subequal, apex obtuse, ribs green, intercostal areas pale green to whitish.
Capsules	6–10(–13) mm.	(5–)6–11 mm, indehiscent.
Anthers	included, ciliate.	(distal pair) nearly exserted, glabrous.
Stigmas	included, lobes unequal, abaxial 1.5 times adaxial.	exserted, lobes unequal, abaxial 3–4 times adaxial.
2n	= 16.

	Diplacus layneae	Diplacus brandegeei
Phenology	Flowering May–Aug.	Flowering Mar–Apr(–May).
Habitat	Road banks, serpentine, granitic sand, red clay, lava beds and volcanic soils, openings in chaparral, shallow dry streambeds or stream banks, burned or otherwise disturbed open areas.	Rocky, brushy slopes.
Elevation	(100–)400–2400 m. ((300–)1300–7900 ft.)	100–300 m. (300–1000 ft.)
Distribution	CA [WildflowerSearch map]	CA; Mexico (Baja California) [BONAP county map]
Discussion	D. M. Thompson (2005) observed that two forms of Diplacus layneae co-occur from the Yosemite National Park area southward; one of these is recognized here as D. graniticola. (Discussion copyrighted by Flora of North America; reprinted with permission.)	Diplacus brandegeei is known only from Santa Catalina (Los Angeles County), Santa Cruz (Santa Barbara County), and Guadalupe (Baja California) islands. The earliest name for the species, Mimulus latifolius A. Gray, was first transferred to Diplacus as D. latifolius (A. Gray) G. L. Nesom, but that name is a later homonym of D. latifolius Nuttall. (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 17, p. 437.	FNA vol. 17, p. 446.
Parent taxa	Phrymaceae > Diplacus	Phrymaceae > Diplacus
Sibling taxa	D. angustatus, D. aridus, D. aurantiacus, D. bicolor, D. bigelovii, D. bolanderi, D. brandegeei, D. brevipes, D. calycinus, D. cascadensis, D. clevelandii, D. clivicola, D. compactus, D. congdonii, D. constrictus, D. cusickii, D. cusickioides, D. deschutesensis, D. douglasii, D. fremontii, D. grandiflorus, D. graniticola, D. jepsonii, D. johnstonii, D. kelloggii, D. leptaleus, D. linearis, D. longiflorus, D. mephiticus, D. mohavensis, D. nanus, D. ovatus, D. parryi, D. parviflorus, D. pictus, D. pulchellus, D. puniceus, D. pygmaeus, D. rattanii, D. rupicola, D. rutilus, D. thompsonii, D. torreyi, D. traskiae, D. tricolor, D. vandenbergensis, D. viscidus	D. angustatus, D. aridus, D. aurantiacus, D. bicolor, D. bigelovii, D. bolanderi, D. brevipes, D. calycinus, D. cascadensis, D. clevelandii, D. clivicola, D. compactus, D. congdonii, D. constrictus, D. cusickii, D. cusickioides, D. deschutesensis, D. douglasii, D. fremontii, D. grandiflorus, D. graniticola, D. jepsonii, D. johnstonii, D. kelloggii, D. layneae, D. leptaleus, D. linearis, D. longiflorus, D. mephiticus, D. mohavensis, D. nanus, D. ovatus, D. parryi, D. parviflorus, D. pictus, D. pulchellus, D. puniceus, D. pygmaeus, D. rattanii, D. rupicola, D. rutilus, D. thompsonii, D. torreyi, D. traskiae, D. tricolor, D. vandenbergensis, D. viscidus
Synonyms	Eunanus layneae, Mimulus brachiatus, M. layneae	Mimulus brandegeei, M. latifolius
Name authority	(Greene) G. L. Nesom: Phytoneuron 2012-39: 29. (2012)	(Pennell) G. L. Nesom: Phytoneuron 2012-54: 1. (2012)
Web links	Local floras: CA Local Web sites: CalFlora, CalPhotos WildflowerSearch iNaturalist (observations) LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	Local floras: CA Local Web sites: CalFlora, CalPhotos WildflowerSearch iNaturalist (observations) LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Diplacus brandegeei

Layne's monkeyflower

Santa Cruz Island monkeyflower, Santa Cruz Island or broadleaf monkeyflower

Habit

Herbs, annual, herbage usually drying dark.

Herbs, annual.

Stems

erect, 30–160(–300) mm, nodes 3–6, glandular-puberulent to glandular-pubescent, hairs 0.2–0.8 mm.

erect, 10–100 mm, glandular-puberulent to glandular-pubescent.

Leaves

usually cauline, relatively even-sized;

petiole weakly delimited;

blade elliptic to narrowly elliptic, oblanceolate, elliptic-oblanceolate, or elliptic-lanceolate, 8–27(–35) × 2–8 mm, margins entire, rarely toothed, plane, apex rounded to obtuse, surfaces: proximals often glabrate, distals glandular-puberulent or glandular-pubescent.

mostly cauline, relatively even-sized;

petiole absent, larger with petiole-like extension;

blade narrowly to broadly ovate to broadly elliptic, (4–)8–39 × (2–)4–23 mm, margins entire or crenate, plane, proximal 1/2 usually ciliate, apex obtuse, surfaces glandular-pubescent.

Pedicels

2–4(–5) mm in fruit.

1.5–4 mm in fruit, not twisting to invert calyx.

Flowers

2 per node, or 1 or 2 per node on 1 plant, chasmogamous.

Styles

glandular-puberulent.

pubescent distally.

Corollas

pinkish or pale to dark magenta or red-purple, each lobe usually with a faint to dark medial line extending 1/2 or less to tip, throat floor yellowish near base, mostly white with red-purple dots near mouth, palate ridges white, tube-throat 10–15 mm, limb (8–)10–16 mm diam., not bilabiate.

throat magenta, golden yellow at base, limb magenta, palate ridges golden yellow, purple-speckled, tube-throat 12–19 mm, limb 5–11 mm diam., bilabiate, abaxial lip smaller than adaxial.

Calyces

symmetrically attached to pedicels, not inflated in fruit, (5–)6–8(–9) mm, glandular-puberulent to glandular-pubescent, tube strongly plicate, lobes triangular, subequal, apex acute, ribs broad, darkened, blackish, thickened, strongly raised, intercostal areas whitish, membranous.

distinctly asymmetrically attached to pedicel, inflated in fruit, 8–13 mm, glandular-pubescent, lobes subequal, apex obtuse, ribs green, intercostal areas pale green to whitish.

Capsules

6–10(–13) mm.

(5–)6–11 mm, indehiscent.

Anthers

included, ciliate.

(distal pair) nearly exserted, glabrous.

Stigmas

included, lobes unequal, abaxial 1.5 times adaxial.

exserted, lobes unequal, abaxial 3–4 times adaxial.

= 16.

Diplacus layneae

Diplacus brandegeei

Phenology

Flowering May–Aug.

Flowering Mar–Apr(–May).

Habitat

Road banks, serpentine, granitic sand, red clay, lava beds and volcanic soils, openings in chaparral, shallow dry streambeds or stream banks, burned or otherwise disturbed open areas.

Rocky, brushy slopes.

Elevation

(100–)400–2400 m. ((300–)1300–7900 ft.)

100–300 m. (300–1000 ft.)

Distribution

[WildflowerSearch map]

CA; Mexico (Baja California)

[BONAP county map]

Discussion

D. M. Thompson (2005) observed that two forms of Diplacus layneae co-occur from the Yosemite National Park area southward; one of these is recognized here as D. graniticola.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Diplacus brandegeei is known only from Santa Catalina (Los Angeles County), Santa Cruz (Santa Barbara County), and Guadalupe (Baja California) islands. The earliest name for the species, Mimulus latifolius A. Gray, was first transferred to Diplacus as D. latifolius (A. Gray) G. L. Nesom, but that name is a later homonym of D. latifolius Nuttall.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 17, p. 437.

FNA vol. 17, p. 446.

Parent taxa

Phrymaceae > Diplacus

Sibling taxa

D. angustatus, D. aridus, D. aurantiacus, D. bicolor, D. bigelovii, D. bolanderi, D. brandegeei, D. brevipes, D. calycinus, D. cascadensis, D. clevelandii, D. clivicola, D. compactus, D. congdonii, D. constrictus, D. cusickii, D. cusickioides, D. deschutesensis, D. douglasii, D. fremontii, D. grandiflorus, D. graniticola, D. jepsonii, D. johnstonii, D. kelloggii, D. leptaleus, D. linearis, D. longiflorus, D. mephiticus, D. mohavensis, D. nanus, D. ovatus, D. parryi, D. parviflorus, D. pictus, D. pulchellus, D. puniceus, D. pygmaeus, D. rattanii, D. rupicola, D. rutilus, D. thompsonii, D. torreyi, D. traskiae, D. tricolor, D. vandenbergensis, D. viscidus

D. angustatus, D. aridus, D. aurantiacus, D. bicolor, D. bigelovii, D. bolanderi, D. brevipes, D. calycinus, D. cascadensis, D. clevelandii, D. clivicola, D. compactus, D. congdonii, D. constrictus, D. cusickii, D. cusickioides, D. deschutesensis, D. douglasii, D. fremontii, D. grandiflorus, D. graniticola, D. jepsonii, D. johnstonii, D. kelloggii, D. layneae, D. leptaleus, D. linearis, D. longiflorus, D. mephiticus, D. mohavensis, D. nanus, D. ovatus, D. parryi, D. parviflorus, D. pictus, D. pulchellus, D. puniceus, D. pygmaeus, D. rattanii, D. rupicola, D. rutilus, D. thompsonii, D. torreyi, D. traskiae, D. tricolor, D. vandenbergensis, D. viscidus

Synonyms

Eunanus layneae, Mimulus brachiatus, M. layneae

Mimulus brandegeei, M. latifolius

Name authority

(Greene) G. L. Nesom: Phytoneuron 2012-39: 29. (2012)

(Pennell) G. L. Nesom: Phytoneuron 2012-54: 1. (2012)

Web links

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

Choose a data source: Flora of North America

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Flora of North America species comparison

Diplacus layneae

Diplacus brandegeei

Diplacus layneae

Diplacus brandegeei