Diplacus layneae |
Diplacus bicolor |
|
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Layne's monkeyflower |
harlequin monkeyflower, Whitney's monkeyflower |
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Habit | Herbs, annual, herbage usually drying dark. | Herbs, annual, herbage usually drying dark. |
Stems | erect, 30–160(–300) mm, nodes 3–6, glandular-puberulent to glandular-pubescent, hairs 0.2–0.8 mm. |
erect or ascending, 10–140(–220) mm, glandular-puberulent to glandular-pubescent. |
Leaves | usually cauline, relatively even-sized; petiole weakly delimited; blade elliptic to narrowly elliptic, oblanceolate, elliptic-oblanceolate, or elliptic-lanceolate, 8–27(–35) × 2–8 mm, margins entire, rarely toothed, plane, apex rounded to obtuse, surfaces: proximals often glabrate, distals glandular-puberulent or glandular-pubescent. |
usually cauline, relatively even-sized; petiole absent; blade narrowly elliptic to narrowly oblanceolate, linear-lanceolate, or linear, (4–)7–23(–34) × 1–5(–8) mm, margins entire, plane, not ciliate, apex acute, surfaces glandular-puberulent. |
Pedicels | 2–4(–5) mm in fruit. |
1–3(–4) mm in fruit. |
Flowers | 2 per node, or 1 or 2 per node on 1 plant, chasmogamous. |
2 per node, or 1 or 2 per node on 1 plant, chasmogamous. |
Styles | glandular-puberulent. |
glandular-puberulent. |
Corollas | pinkish or pale to dark magenta or red-purple, each lobe usually with a faint to dark medial line extending 1/2 or less to tip, throat floor yellowish near base, mostly white with red-purple dots near mouth, palate ridges white, tube-throat 10–15 mm, limb (8–)10–16 mm diam., not bilabiate. |
(a) magenta, darkening toward mouth and within tube, often nearly obscuring dark longitudinal stripes within, throat floor or at least palate ridges yellow, or (b) yellow, usually with maroon stripes in throat extending onto lobe bases, sometimes with maroon blotches on adaxial lateral walls of throat, colored palate ridges ending in throat, tube-throat (10–)13–18(–20) mm, limb 10–19 mm diam., not bilabiate. |
Calyces | symmetrically attached to pedicels, not inflated in fruit, (5–)6–8(–9) mm, glandular-puberulent to glandular-pubescent, tube strongly plicate, lobes triangular, subequal, apex acute, ribs broad, darkened, blackish, thickened, strongly raised, intercostal areas whitish, membranous. |
symmetrically attached to pedicels, not inflated in fruit, (3–)4–8(–10) mm, glandular-puberulent, tube not strongly plicate, lobes triangular, subequal, apex acute-apiculate, ribs narrow, darkened, purplish, thin, not strongly raised, intercostal areas pale green. |
Capsules | 6–10(–13) mm. |
(4.5–)6–10(–13) mm. |
Anthers | included, ciliate. |
included, ciliate. |
Stigmas | included, lobes unequal, abaxial 1.5 times adaxial. |
included, lobes equal. |
2n | = 16. |
= 16. |
Diplacus layneae |
Diplacus bicolor |
|
Phenology | Flowering May–Aug. | Flowering May–Sep. |
Habitat | Road banks, serpentine, granitic sand, red clay, lava beds and volcanic soils, openings in chaparral, shallow dry streambeds or stream banks, burned or otherwise disturbed open areas. | Disturbed areas, water runoff areas, granitic soils, edges of granite outcrops. |
Elevation | (100–)400–2400 m. ((300–)1300–7900 ft.) | (1200–)1500–3300 m. ((3900–)4900–10800 ft.) |
Distribution |
CA
|
CA |
Discussion | D. M. Thompson (2005) observed that two forms of Diplacus layneae co-occur from the Yosemite National Park area southward; one of these is recognized here as D. graniticola. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Diplacus bicolor is endemic to Fresno, Kern, and Tulare counties in the southern Sierra Nevada. The combination D. whitneyi (A. Gray) G. L. Nesom is illegitimate, as Mimulus whitneyi A. Gray 1886, is a replacement name based on Eunanus bicolor A. Gray 1868. Diplacus bicolor, D. bigelovii, D. constrictus, D. graniticola, D. layneae, and D. thompsonii appear to be closely related species, sometimes intergrading where sympatric. Plants of each species often produce flowers at all nodes and have dark magenta corollas with nearly regular to weakly bilabiate limbs and villous vestiture. Specimens of each species dry to a dark color. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 437. | FNA vol. 17, p. 437. |
Parent taxa | Phrymaceae > Diplacus | Phrymaceae > Diplacus |
Sibling taxa | ||
Synonyms | Eunanus layneae, Mimulus brachiatus, M. layneae | Eunanus bicolor, M. nanus var. bicolor, M. whitneyi |
Name authority | (Greene) G. L. Nesom: Phytoneuron 2012-39: 29. (2012) | (A. Gray) Hrusa: Phytoneuron 2014-17: 1. (2014) |
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