Flora of North America species comparison

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Diplacus johnstonii

Diplacus vandenbergensis

Johnston's monkeyflower

Vandenberg monkeyflower
Habit Herbs, annual. Herbs, annual.
Stems

erect, (10–)30–200(–300) mm, densely glandular-puberulent.

erect, 10–200(–240) mm, glandular-puberulent or glandular-pubescent.
Leaves

basal and cauline, gradually reduced distally;

petiole absent, bases of larger leaves often with petiole-like extensions;

blade obovate or oblanceolate, sometimes elliptic, (4.5–)7–25(–32) × 2–12(–15) mm, margins entire, plane, apex rounded to acute, surfaces densely glandular-puberulent.

basal and cauline, basal in rosette, cauline reduced distally;

petiole absent;

blade narrowly elliptic, sometimes obovate to oblanceolate, 2–30(–55) × 1–10(–16) mm, margins entire, sometimes crenate to serrate, plane, not ciliate, apex rounded to acute, surfaces: proximals glabrous, distals glandular-puberulent or glandular-pubescent.
Pedicels

1–4(–5) mm in fruit.

1–4(–7 on proximal) mm in fruit.
Flowers

2 per node, or 1 or 2 per node on 1 plant, chasmogamous.

1 per node, chasmogamous.
Styles

densely glandular-puberulent distally.

minutely puberulent.
Corollas

magenta, darker and more reddish in throat and, often, along narrow radiating lines extending from throat onto midveins of lobes, throat usually with a large dark spot on each side of mouth on lateral walls, palate ridges and throat floor yellow with reddish spots, tube-throat 9–15 mm, limb 10–15 mm diam., not bilabiate.

yellow with reddish brown spots near mouth, palate ridges yellow, throat floor puberulent, tube-throat (9–)10–23 mm, limb 8–26 mm diam., not bilabiate.
Calyces

symmetrically attached to pedicels, not inflated in fruit, (6–)7–11 mm, glandular-puberulent, lobes unequal, apex acute to acuminate, ribs inconspicuous, intercostal areas reddish.

inflated in fruit, 5–14 mm, glandular-puberulent to glandular-pubescent or ribs almost tomentose and viscid, lobes subequal, apex rounded and apiculate or acute, intercostal areas white.
Capsules

7–12 mm.

6.5–13(–14) mm.
Anthers

included, glabrous.

included, glabrous, rarely minutely puberulent.
Stigmas

exserted or at opening of corolla tube-throat, lobes equal.

included, lobes equal.
2n

= 16.

= 16.

Diplacus johnstonii

Diplacus vandenbergensis
Phenology Flowering May–Aug. Flowering Apr–Jun.
Habitat Steep, unstable scree slides, talus slopes, gravel slides, cracks in granite cliffs, ridges, washes, steep sand and gravel slopes, canyon bottoms, gravelly road banks, recent burns, desert scrub, chaparral, juniper, pinyon-juniper, lodgepole pine, yellow pine, Jeffrey pine, and Jeffrey pine-western white pine-fir woodlands. Sandy open or disturbed areas, among shrubs.
Elevation (1000–)1300–2900 m. ((3300–)4300–9500 ft.) 70–150 m. (200–500 ft.)
Distribution
from FNA
CA
[WildflowerSearch map]
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
Discussion

Populations of Diplacus johnstonii occur in Los Angeles, Riverside, San Bernardino, Santa Barbara, and Ventura counties. The populations in northwestern Los Angeles, Santa Barbara, and Ventura counties were noted by D. M. Thompson (2005) to be intermediate between D. constrictus and D. johnstonii.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Diplacus vandenbergensis is endemic to Santa Barbara County, mostly on the north side of the city of Lompoc (La Purisima Mission State Historic Park; also on and near Vandenberg Air Force Base) plus one other locality about 10 kilometers farther west (Santa Ynez Valley, 8 miles west of Buellton). Thompson described Mimulus fremontii var. vandenbergensis as a geographically distinct variant of M. fremontii. The discontinuous morphological difference, allopatric/parapatric geographical distribution, and lack of intergrading populations support recognition of the plants with yellow corollas at specific rank. Plants with yellow corollas are found in D. bicolor, D. mephiticus, and D. parryi, but these morphs are found through most of the geographic ranges of these taxa and sometimes are intermixed in populations.

(Discussion copyrighted by Flora of North America; reprinted with permission.)
Source FNA vol. 17, p. 433. FNA vol. 17, p. 432.
Parent taxa Phrymaceae > Diplacus Phrymaceae > Diplacus
Sibling taxa
D. angustatus, D. aridus, D. aurantiacus, D. bicolor, D. bigelovii, D. bolanderi, D. brandegeei, D. brevipes, D. calycinus, D. cascadensis, D. clevelandii, D. clivicola, D. compactus, D. congdonii, D. constrictus, D. cusickii, D. cusickioides, D. deschutesensis, D. douglasii, D. fremontii, D. grandiflorus, D. graniticola, D. jepsonii, D. kelloggii, D. layneae, D. leptaleus, D. linearis, D. longiflorus, D. mephiticus, D. mohavensis, D. nanus, D. ovatus, D. parryi, D. parviflorus, D. pictus, D. pulchellus, D. puniceus, D. pygmaeus, D. rattanii, D. rupicola, D. rutilus, D. thompsonii, D. torreyi, D. traskiae, D. tricolor, D. vandenbergensis, D. viscidus
D. angustatus, D. aridus, D. aurantiacus, D. bicolor, D. bigelovii, D. bolanderi, D. brandegeei, D. brevipes, D. calycinus, D. cascadensis, D. clevelandii, D. clivicola, D. compactus, D. congdonii, D. constrictus, D. cusickii, D. cusickioides, D. deschutesensis, D. douglasii, D. fremontii, D. grandiflorus, D. graniticola, D. jepsonii, D. johnstonii, D. kelloggii, D. layneae, D. leptaleus, D. linearis, D. longiflorus, D. mephiticus, D. mohavensis, D. nanus, D. ovatus, D. parryi, D. parviflorus, D. pictus, D. pulchellus, D. puniceus, D. pygmaeus, D. rattanii, D. rupicola, D. rutilus, D. thompsonii, D. torreyi, D. traskiae, D. tricolor, D. viscidus
Synonyms Mimulus johnstonii Mimulus fremontii var. vandenbergensis
Name authority (A. L. Grant) G. L. Nesom: Phytoneuron 2012-39: 29. (2012) (D. M. Thompson) G. L. Nesom: Phytoneuron 2012-47: 2. (2012)
Web links 
Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).



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