FNA: Diplacus johnstonii vs. Diplacus clivicola

	Diplacus johnstonii	Diplacus clivicola
	Johnston's monkeyflower	bank monkey-flower, north Idaho monkeyflower, slope monkeyflower
Habit	Herbs, annual.	Herbs, annual.
Stems	erect, (10–)30–200(–300) mm, densely glandular-puberulent.	erect, (10–)20–180 mm, glandular-puberulent to short glandular-villous.
Leaves	basal and cauline, gradually reduced distally; petiole absent, bases of larger leaves often with petiole-like extensions; blade obovate or oblanceolate, sometimes elliptic, (4.5–)7–25(–32) × 2–12(–15) mm, margins entire, plane, apex rounded to acute, surfaces densely glandular-puberulent.	usually cauline, relatively even-sized; petiole absent, base sometimes petiole-like; blade narrowly elliptic, sometimes broadly elliptic to obovate or oblanceolate, (2.5–)6–20(–26) × (1–)2–10(–14) mm, margins crenate to serrulate or entire, plane, apex rounded or acute, surfaces glandular-puberulent.
Pedicels	1–4(–5) mm in fruit.	2–7(–10) mm in fruit.
Flowers	2 per node, or 1 or 2 per node on 1 plant, chasmogamous.	2 per node, or 1 or 2 per node on 1 plant, chasmogamous.
Styles	densely glandular-puberulent distally.	glandular-puberulent distally.
Corollas	magenta, darker and more reddish in throat and, often, along narrow radiating lines extending from throat onto midveins of lobes, throat usually with a large dark spot on each side of mouth on lateral walls, palate ridges and throat floor yellow with reddish spots, tube-throat 9–15 mm, limb 10–15 mm diam., not bilabiate.	rose pink to purplish, limb often pale, especially abaxial lip, abaxial lip often purple-dotted near base, markings often coalescing and forming broken lines radiating toward each lobe, tube yellow, palate ridges yellow with magenta speckling, confluent and extending onto abaxial lip base, tube-throat (8–)11–12 mm, limb 7–12 mm diam., bilabiate.
Calyces	symmetrically attached to pedicels, not inflated in fruit, (6–)7–11 mm, glandular-puberulent, lobes unequal, apex acute to acuminate, ribs inconspicuous, intercostal areas reddish.	symmetrically attached to pedicels, not inflated in fruit, (5–)7–8 mm, glandular-puberulent, lobes subequal, apex acute, ribs green, intercostal areas whitish.
Capsules	7–12 mm.	8–13 mm.
Anthers	included, glabrous.	included, ciliate.
Stigmas	exserted or at opening of corolla tube-throat, lobes equal.	exserted, lobes subequal.
2n	= 16.	= 16.

	Diplacus johnstonii	Diplacus clivicola
Phenology	Flowering May–Aug.	Flowering May–Aug.
Habitat	Steep, unstable scree slides, talus slopes, gravel slides, cracks in granite cliffs, ridges, washes, steep sand and gravel slopes, canyon bottoms, gravelly road banks, recent burns, desert scrub, chaparral, juniper, pinyon-juniper, lodgepole pine, yellow pine, Jeffrey pine, and Jeffrey pine-western white pine-fir woodlands.	Bluffs, disturbed slopes, well-developed loam soils, vegetation openings.
Elevation	(1000–)1300–2900 m. ((3300–)4300–9500 ft.)	500–1200(–2000) m. (1600–3900(–6600) ft.)
Distribution	CA [WildflowerSearch map]	ID; MT; OR [WildflowerSearch map]
Discussion	Populations of Diplacus johnstonii occur in Los Angeles, Riverside, San Bernardino, Santa Barbara, and Ventura counties. The populations in northwestern Los Angeles, Santa Barbara, and Ventura counties were noted by D. M. Thompson (2005) to be intermediate between D. constrictus and D. johnstonii. (Discussion copyrighted by Flora of North America; reprinted with permission.)	Diplacus clivicola is known from northern Idaho and immediately adjacent Montana and Oregon. It is similar to typical D. nanus in its strongly bilabiate corollas; it differs in its slightly toothed leaf blade margins, relatively long pedicels, and calyces with cuneate bases. (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 17, p. 433.	FNA vol. 17, p. 438.
Parent taxa	Phrymaceae > Diplacus	Phrymaceae > Diplacus
Sibling taxa	D. angustatus, D. aridus, D. aurantiacus, D. bicolor, D. bigelovii, D. bolanderi, D. brandegeei, D. brevipes, D. calycinus, D. cascadensis, D. clevelandii, D. clivicola, D. compactus, D. congdonii, D. constrictus, D. cusickii, D. cusickioides, D. deschutesensis, D. douglasii, D. fremontii, D. grandiflorus, D. graniticola, D. jepsonii, D. kelloggii, D. layneae, D. leptaleus, D. linearis, D. longiflorus, D. mephiticus, D. mohavensis, D. nanus, D. ovatus, D. parryi, D. parviflorus, D. pictus, D. pulchellus, D. puniceus, D. pygmaeus, D. rattanii, D. rupicola, D. rutilus, D. thompsonii, D. torreyi, D. traskiae, D. tricolor, D. vandenbergensis, D. viscidus	D. angustatus, D. aridus, D. aurantiacus, D. bicolor, D. bigelovii, D. bolanderi, D. brandegeei, D. brevipes, D. calycinus, D. cascadensis, D. clevelandii, D. compactus, D. congdonii, D. constrictus, D. cusickii, D. cusickioides, D. deschutesensis, D. douglasii, D. fremontii, D. grandiflorus, D. graniticola, D. jepsonii, D. johnstonii, D. kelloggii, D. layneae, D. leptaleus, D. linearis, D. longiflorus, D. mephiticus, D. mohavensis, D. nanus, D. ovatus, D. parryi, D. parviflorus, D. pictus, D. pulchellus, D. puniceus, D. pygmaeus, D. rattanii, D. rupicola, D. rutilus, D. thompsonii, D. torreyi, D. traskiae, D. tricolor, D. vandenbergensis, D. viscidus
Synonyms	Mimulus johnstonii	Mimulus clivicola
Name authority	(A. L. Grant) G. L. Nesom: Phytoneuron 2012-39: 29. (2012)	(Greenman) G. L. Nesom: Phytoneuron 2012-39: 28. (2012)
Web links	Local floras: CA Local Web sites: CalFlora, CalPhotos WildflowerSearch iNaturalist (observations) LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	Local floras: OR Local Web sites: Flora NW, PNW Herbaria WildflowerSearch iNaturalist (observations) LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Diplacus clivicola

Johnston's monkeyflower

bank monkey-flower, north Idaho monkeyflower, slope monkeyflower

Habit

Herbs, annual.

Stems

erect, (10–)30–200(–300) mm, densely glandular-puberulent.

erect, (10–)20–180 mm, glandular-puberulent to short glandular-villous.

Leaves

basal and cauline, gradually reduced distally;

petiole absent, bases of larger leaves often with petiole-like extensions;

blade obovate or oblanceolate, sometimes elliptic, (4.5–)7–25(–32) × 2–12(–15) mm, margins entire, plane, apex rounded to acute, surfaces densely glandular-puberulent.

usually cauline, relatively even-sized;

petiole absent, base sometimes petiole-like;

blade narrowly elliptic, sometimes broadly elliptic to obovate or oblanceolate, (2.5–)6–20(–26) × (1–)2–10(–14) mm, margins crenate to serrulate or entire, plane, apex rounded or acute, surfaces glandular-puberulent.

Pedicels

1–4(–5) mm in fruit.

2–7(–10) mm in fruit.

Flowers

2 per node, or 1 or 2 per node on 1 plant, chasmogamous.

Styles

densely glandular-puberulent distally.

glandular-puberulent distally.

Corollas

magenta, darker and more reddish in throat and, often, along narrow radiating lines extending from throat onto midveins of lobes, throat usually with a large dark spot on each side of mouth on lateral walls, palate ridges and throat floor yellow with reddish spots, tube-throat 9–15 mm, limb 10–15 mm diam., not bilabiate.

rose pink to purplish, limb often pale, especially abaxial lip, abaxial lip often purple-dotted near base, markings often coalescing and forming broken lines radiating toward each lobe, tube yellow, palate ridges yellow with magenta speckling, confluent and extending onto abaxial lip base, tube-throat (8–)11–12 mm, limb 7–12 mm diam., bilabiate.

Calyces

symmetrically attached to pedicels, not inflated in fruit, (6–)7–11 mm, glandular-puberulent, lobes unequal, apex acute to acuminate, ribs inconspicuous, intercostal areas reddish.

symmetrically attached to pedicels, not inflated in fruit, (5–)7–8 mm, glandular-puberulent, lobes subequal, apex acute, ribs green, intercostal areas whitish.

Capsules

7–12 mm.

8–13 mm.

Anthers

included, glabrous.

included, ciliate.

Stigmas

exserted or at opening of corolla tube-throat, lobes equal.

exserted, lobes subequal.

= 16.

Diplacus johnstonii

Diplacus clivicola

Phenology

Flowering May–Aug.

Habitat

Steep, unstable scree slides, talus slopes, gravel slides, cracks in granite cliffs, ridges, washes, steep sand and gravel slopes, canyon bottoms, gravelly road banks, recent burns, desert scrub, chaparral, juniper, pinyon-juniper, lodgepole pine, yellow pine, Jeffrey pine, and Jeffrey pine-western white pine-fir woodlands.

Bluffs, disturbed slopes, well-developed loam soils, vegetation openings.

Elevation

(1000–)1300–2900 m. ((3300–)4300–9500 ft.)

500–1200(–2000) m. (1600–3900(–6600) ft.)

Distribution

[WildflowerSearch map]

ID; MT; OR

[WildflowerSearch map]

Discussion

Populations of Diplacus johnstonii occur in Los Angeles, Riverside, San Bernardino, Santa Barbara, and Ventura counties. The populations in northwestern Los Angeles, Santa Barbara, and Ventura counties were noted by D. M. Thompson (2005) to be intermediate between D. constrictus and D. johnstonii.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Diplacus clivicola is known from northern Idaho and immediately adjacent Montana and Oregon. It is similar to typical D. nanus in its strongly bilabiate corollas; it differs in its slightly toothed leaf blade margins, relatively long pedicels, and calyces with cuneate bases.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 17, p. 433.

FNA vol. 17, p. 438.

Parent taxa

Phrymaceae > Diplacus

Sibling taxa

D. angustatus, D. aridus, D. aurantiacus, D. bicolor, D. bigelovii, D. bolanderi, D. brandegeei, D. brevipes, D. calycinus, D. cascadensis, D. clevelandii, D. clivicola, D. compactus, D. congdonii, D. constrictus, D. cusickii, D. cusickioides, D. deschutesensis, D. douglasii, D. fremontii, D. grandiflorus, D. graniticola, D. jepsonii, D. kelloggii, D. layneae, D. leptaleus, D. linearis, D. longiflorus, D. mephiticus, D. mohavensis, D. nanus, D. ovatus, D. parryi, D. parviflorus, D. pictus, D. pulchellus, D. puniceus, D. pygmaeus, D. rattanii, D. rupicola, D. rutilus, D. thompsonii, D. torreyi, D. traskiae, D. tricolor, D. vandenbergensis, D. viscidus

D. angustatus, D. aridus, D. aurantiacus, D. bicolor, D. bigelovii, D. bolanderi, D. brandegeei, D. brevipes, D. calycinus, D. cascadensis, D. clevelandii, D. compactus, D. congdonii, D. constrictus, D. cusickii, D. cusickioides, D. deschutesensis, D. douglasii, D. fremontii, D. grandiflorus, D. graniticola, D. jepsonii, D. johnstonii, D. kelloggii, D. layneae, D. leptaleus, D. linearis, D. longiflorus, D. mephiticus, D. mohavensis, D. nanus, D. ovatus, D. parryi, D. parviflorus, D. pictus, D. pulchellus, D. puniceus, D. pygmaeus, D. rattanii, D. rupicola, D. rutilus, D. thompsonii, D. torreyi, D. traskiae, D. tricolor, D. vandenbergensis, D. viscidus

Synonyms

Mimulus johnstonii

Mimulus clivicola

Name authority

(A. L. Grant) G. L. Nesom: Phytoneuron 2012-39: 29. (2012)

(Greenman) G. L. Nesom: Phytoneuron 2012-39: 28. (2012)

Web links

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

Choose a data source: Flora of North America, OregonFlora

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Flora of North America species comparison

Diplacus johnstonii

Diplacus clivicola

Diplacus johnstonii

Diplacus clivicola