FNA: Diplacus johnstonii vs. Diplacus bolanderi

	Diplacus johnstonii	Diplacus bolanderi
	Johnston's monkeyflower	Bolander's monkeyflower
Habit	Herbs, annual.	Herbs, annual.
Stems	erect, (10–)30–200(–300) mm, densely glandular-puberulent.	erect, 20–900(–1200) mm, usually glandular-pubescent and viscid.
Leaves	basal and cauline, gradually reduced distally; petiole absent, bases of larger leaves often with petiole-like extensions; blade obovate or oblanceolate, sometimes elliptic, (4.5–)7–25(–32) × 2–12(–15) mm, margins entire, plane, apex rounded to acute, surfaces densely glandular-puberulent.	basal and cauline, basal in rosette, cauline gradually reduced distally; petiole present proximally, usually absent distally; blade ovate, obovate, elliptic, or oblanceolate, 5–60(–85) × 1.5–25(–32) mm, margins entire or largest irregularly serrate, plane, apex rounded to obtuse, surfaces: proximals nearly glabrous, distals glandular-pubescent.
Pedicels	1–4(–5) mm in fruit.	2–5(–8) mm in fruit.
Flowers	2 per node, or 1 or 2 per node on 1 plant, chasmogamous.	2 per node, or 1 or 2 per node on 1 plant (1 per node on depauperate plants), chasmogamous.
Styles	densely glandular-puberulent distally.	glandular-puberulent.
Corollas	magenta, darker and more reddish in throat and, often, along narrow radiating lines extending from throat onto midveins of lobes, throat usually with a large dark spot on each side of mouth on lateral walls, palate ridges and throat floor yellow with reddish spots, tube-throat 9–15 mm, limb 10–15 mm diam., not bilabiate.	magenta, throat often darkening near floor, without dark spots, palate ridges or floor white, usually magenta-speckled, tube-throat 10–30 mm, limb 6–20 mm diam., not bilabiate.
Calyces	symmetrically attached to pedicels, not inflated in fruit, (6–)7–11 mm, glandular-puberulent, lobes unequal, apex acute to acuminate, ribs inconspicuous, intercostal areas reddish.	symmetrically attached to pedicels, inflated in fruit, (7–)9–25(–27) mm, glandular-pubescent and viscid, lobes often spreading in fruit, unequal if calyx large, apex acute to attenuate, ribs prominent, purplish, intercostal areas white.
Capsules	7–12 mm.	8–20 mm.
Anthers	included, glabrous.	included, glabrous.
Stigmas	exserted or at opening of corolla tube-throat, lobes equal.	exserted, lobes unequal, abaxial 2–3 times adaxial.
2n	= 16.	= 16.

	Diplacus johnstonii	Diplacus bolanderi
Phenology	Flowering May–Aug.	Flowering Apr–Jul.
Habitat	Steep, unstable scree slides, talus slopes, gravel slides, cracks in granite cliffs, ridges, washes, steep sand and gravel slopes, canyon bottoms, gravelly road banks, recent burns, desert scrub, chaparral, juniper, pinyon-juniper, lodgepole pine, yellow pine, Jeffrey pine, and Jeffrey pine-western white pine-fir woodlands.	Granite outcrops, slopes with decomposed granite soils.
Elevation	(1000–)1300–2900 m. ((3300–)4300–9500 ft.)	(100–)300–1700(–2000) m. ((300–)1000–5600(–6600) ft.)
Distribution	CA [WildflowerSearch map]	CA; OR [WildflowerSearch map]
Discussion	Populations of Diplacus johnstonii occur in Los Angeles, Riverside, San Bernardino, Santa Barbara, and Ventura counties. The populations in northwestern Los Angeles, Santa Barbara, and Ventura counties were noted by D. M. Thompson (2005) to be intermediate between D. constrictus and D. johnstonii. (Discussion copyrighted by Flora of North America; reprinted with permission.)	Diplacus bolanderi is scattered and uncommon through most of California (with concentrations in the southern North Coast Ranges and Central Sierra) and apparently is disjunct in Jackson County, Oregon. Diplacus bolanderi is highly variable in height, leaf size, corolla size, and calyx size. Identity is unmistakable in larger plants; the inflated calyx with lobes strongly unequal in length usually is helpful in identification of smaller plants. (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 17, p. 433.	FNA vol. 17, p. 434.
Parent taxa	Phrymaceae > Diplacus	Phrymaceae > Diplacus
Sibling taxa	D. angustatus, D. aridus, D. aurantiacus, D. bicolor, D. bigelovii, D. bolanderi, D. brandegeei, D. brevipes, D. calycinus, D. cascadensis, D. clevelandii, D. clivicola, D. compactus, D. congdonii, D. constrictus, D. cusickii, D. cusickioides, D. deschutesensis, D. douglasii, D. fremontii, D. grandiflorus, D. graniticola, D. jepsonii, D. kelloggii, D. layneae, D. leptaleus, D. linearis, D. longiflorus, D. mephiticus, D. mohavensis, D. nanus, D. ovatus, D. parryi, D. parviflorus, D. pictus, D. pulchellus, D. puniceus, D. pygmaeus, D. rattanii, D. rupicola, D. rutilus, D. thompsonii, D. torreyi, D. traskiae, D. tricolor, D. vandenbergensis, D. viscidus	D. angustatus, D. aridus, D. aurantiacus, D. bicolor, D. bigelovii, D. brandegeei, D. brevipes, D. calycinus, D. cascadensis, D. clevelandii, D. clivicola, D. compactus, D. congdonii, D. constrictus, D. cusickii, D. cusickioides, D. deschutesensis, D. douglasii, D. fremontii, D. grandiflorus, D. graniticola, D. jepsonii, D. johnstonii, D. kelloggii, D. layneae, D. leptaleus, D. linearis, D. longiflorus, D. mephiticus, D. mohavensis, D. nanus, D. ovatus, D. parryi, D. parviflorus, D. pictus, D. pulchellus, D. puniceus, D. pygmaeus, D. rattanii, D. rupicola, D. rutilus, D. thompsonii, D. torreyi, D. traskiae, D. tricolor, D. vandenbergensis, D. viscidus
Synonyms	Mimulus johnstonii	Mimulus bolanderi, M. platylaemus
Name authority	(A. L. Grant) G. L. Nesom: Phytoneuron 2012-39: 29. (2012)	(A. Gray) G. L. Nesom: Phytoneuron 2012-39: 28. (2012)
Web links	Local floras: CA Local Web sites: CalFlora, CalPhotos WildflowerSearch iNaturalist (observations) LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	Local floras: CA, OR Local Web sites: CalFlora, CalPhotos, Flora NW, PNW Herbaria WildflowerSearch iNaturalist (observations) LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Diplacus bolanderi

Johnston's monkeyflower

Bolander's monkeyflower

Habit

Herbs, annual.

Stems

erect, (10–)30–200(–300) mm, densely glandular-puberulent.

erect, 20–900(–1200) mm, usually glandular-pubescent and viscid.

Leaves

basal and cauline, gradually reduced distally;

petiole absent, bases of larger leaves often with petiole-like extensions;

blade obovate or oblanceolate, sometimes elliptic, (4.5–)7–25(–32) × 2–12(–15) mm, margins entire, plane, apex rounded to acute, surfaces densely glandular-puberulent.

basal and cauline, basal in rosette, cauline gradually reduced distally;

petiole present proximally, usually absent distally;

blade ovate, obovate, elliptic, or oblanceolate, 5–60(–85) × 1.5–25(–32) mm, margins entire or largest irregularly serrate, plane, apex rounded to obtuse, surfaces: proximals nearly glabrous, distals glandular-pubescent.

Pedicels

1–4(–5) mm in fruit.

2–5(–8) mm in fruit.

Flowers

2 per node, or 1 or 2 per node on 1 plant, chasmogamous.

2 per node, or 1 or 2 per node on 1 plant (1 per node on depauperate plants), chasmogamous.

Styles

densely glandular-puberulent distally.

glandular-puberulent.

Corollas

magenta, darker and more reddish in throat and, often, along narrow radiating lines extending from throat onto midveins of lobes, throat usually with a large dark spot on each side of mouth on lateral walls, palate ridges and throat floor yellow with reddish spots, tube-throat 9–15 mm, limb 10–15 mm diam., not bilabiate.

magenta, throat often darkening near floor, without dark spots, palate ridges or floor white, usually magenta-speckled, tube-throat 10–30 mm, limb 6–20 mm diam., not bilabiate.

Calyces

symmetrically attached to pedicels, not inflated in fruit, (6–)7–11 mm, glandular-puberulent, lobes unequal, apex acute to acuminate, ribs inconspicuous, intercostal areas reddish.

symmetrically attached to pedicels, inflated in fruit, (7–)9–25(–27) mm, glandular-pubescent and viscid, lobes often spreading in fruit, unequal if calyx large, apex acute to attenuate, ribs prominent, purplish, intercostal areas white.

Capsules

7–12 mm.

8–20 mm.

Anthers

included, glabrous.

Stigmas

exserted or at opening of corolla tube-throat, lobes equal.

exserted, lobes unequal, abaxial 2–3 times adaxial.

= 16.

Diplacus johnstonii

Diplacus bolanderi

Phenology

Flowering May–Aug.

Flowering Apr–Jul.

Habitat

Steep, unstable scree slides, talus slopes, gravel slides, cracks in granite cliffs, ridges, washes, steep sand and gravel slopes, canyon bottoms, gravelly road banks, recent burns, desert scrub, chaparral, juniper, pinyon-juniper, lodgepole pine, yellow pine, Jeffrey pine, and Jeffrey pine-western white pine-fir woodlands.

Granite outcrops, slopes with decomposed granite soils.

Elevation

(1000–)1300–2900 m. ((3300–)4300–9500 ft.)

(100–)300–1700(–2000) m. ((300–)1000–5600(–6600) ft.)

Distribution

[WildflowerSearch map]

CA; OR

[WildflowerSearch map]

Discussion

Populations of Diplacus johnstonii occur in Los Angeles, Riverside, San Bernardino, Santa Barbara, and Ventura counties. The populations in northwestern Los Angeles, Santa Barbara, and Ventura counties were noted by D. M. Thompson (2005) to be intermediate between D. constrictus and D. johnstonii.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Diplacus bolanderi is scattered and uncommon through most of California (with concentrations in the southern North Coast Ranges and Central Sierra) and apparently is disjunct in Jackson County, Oregon.

Diplacus bolanderi is highly variable in height, leaf size, corolla size, and calyx size. Identity is unmistakable in larger plants; the inflated calyx with lobes strongly unequal in length usually is helpful in identification of smaller plants.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 17, p. 433.

FNA vol. 17, p. 434.

Parent taxa

Phrymaceae > Diplacus

Sibling taxa

D. angustatus, D. aridus, D. aurantiacus, D. bicolor, D. bigelovii, D. bolanderi, D. brandegeei, D. brevipes, D. calycinus, D. cascadensis, D. clevelandii, D. clivicola, D. compactus, D. congdonii, D. constrictus, D. cusickii, D. cusickioides, D. deschutesensis, D. douglasii, D. fremontii, D. grandiflorus, D. graniticola, D. jepsonii, D. kelloggii, D. layneae, D. leptaleus, D. linearis, D. longiflorus, D. mephiticus, D. mohavensis, D. nanus, D. ovatus, D. parryi, D. parviflorus, D. pictus, D. pulchellus, D. puniceus, D. pygmaeus, D. rattanii, D. rupicola, D. rutilus, D. thompsonii, D. torreyi, D. traskiae, D. tricolor, D. vandenbergensis, D. viscidus

D. angustatus, D. aridus, D. aurantiacus, D. bicolor, D. bigelovii, D. brandegeei, D. brevipes, D. calycinus, D. cascadensis, D. clevelandii, D. clivicola, D. compactus, D. congdonii, D. constrictus, D. cusickii, D. cusickioides, D. deschutesensis, D. douglasii, D. fremontii, D. grandiflorus, D. graniticola, D. jepsonii, D. johnstonii, D. kelloggii, D. layneae, D. leptaleus, D. linearis, D. longiflorus, D. mephiticus, D. mohavensis, D. nanus, D. ovatus, D. parryi, D. parviflorus, D. pictus, D. pulchellus, D. puniceus, D. pygmaeus, D. rattanii, D. rupicola, D. rutilus, D. thompsonii, D. torreyi, D. traskiae, D. tricolor, D. vandenbergensis, D. viscidus

Synonyms

Mimulus johnstonii

Mimulus bolanderi, M. platylaemus

Name authority

(A. L. Grant) G. L. Nesom: Phytoneuron 2012-39: 29. (2012)

(A. Gray) G. L. Nesom: Phytoneuron 2012-39: 28. (2012)

Web links

Local floras: CA, OR
Local Web sites: CalFlora, CalPhotos, Flora NW, PNW Herbaria
WildflowerSearch
iNaturalist (observations)
LBJ Wildflower Center
SEINet
Plants of the World Online
Encyclopedia of Life
Wikipedia
Google Image Search

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

Choose a data source: Flora of North America, OregonFlora

The species comparison tool is brought to you by:

Flora of North America species comparison

Diplacus johnstonii

Diplacus bolanderi

Diplacus johnstonii

Diplacus bolanderi