Flora of North America species comparison

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Diplacus graniticola

Diplacus vandenbergensis

granite-crack monkeyflower

Vandenberg monkeyflower
Habit Herbs, annual, herbage usually drying dark. Herbs, annual.
Stems

erect, 60–120(–150) mm, nodes 4–15(–20), internodes shorter than leaves, glandular-villous with gland-tipped hairs 1–1.6 mm.

erect, 10–200(–240) mm, glandular-puberulent or glandular-pubescent.
Leaves

usually cauline, relatively even-sized;

petiole weakly delimited;

blade usually lanceolate to ovate-lanceolate, 20–40 × 4–12 mm, margins entire, rarely toothed, plane, apex rounded to obtuse or acute, surfaces: proximals often glabrate abaxially, distals glandular-villous.

basal and cauline, basal in rosette, cauline reduced distally;

petiole absent;

blade narrowly elliptic, sometimes obovate to oblanceolate, 2–30(–55) × 1–10(–16) mm, margins entire, sometimes crenate to serrate, plane, not ciliate, apex rounded to acute, surfaces: proximals glabrous, distals glandular-puberulent or glandular-pubescent.
Pedicels

1–3 mm in fruit.

1–4(–7 on proximal) mm in fruit.
Flowers

2 per node, or 1 or 2 per node on 1 plant, chasmogamous.

1 per node, chasmogamous.
Styles

glandular-puberulent.

minutely puberulent.
Corollas

nearly white or pale lavender to pinkish or pale to dark magenta, each lobe with a dark medial line extending nearly to tip, throat with a dark red or purple splotch at junction of each abaxial lobe and adjacent lateral lobe, throat floor sometimes with 2 adjacent white splotches at lateral lobe bases, palate ridges yellow, tube-throat 15–20 mm, limb 10–16 mm diam., bilabiate.

yellow with reddish brown spots near mouth, palate ridges yellow, throat floor puberulent, tube-throat (9–)10–23 mm, limb 8–26 mm diam., not bilabiate.
Calyces

symmetrically attached to pedicels, not inflated in fruit, 8–12 mm, glandular-villous, tube strongly plicate, lobes triangular, subequal, apex acute, ribs narrow, darkened, blackish, thickened, strongly raised, intercostal areas green to purple, not membranous.

inflated in fruit, 5–14 mm, glandular-puberulent to glandular-pubescent or ribs almost tomentose and viscid, lobes subequal, apex rounded and apiculate or acute, intercostal areas white.
Capsules

6–10 mm.

6.5–13(–14) mm.
Anthers

included, ciliate.

included, glabrous, rarely minutely puberulent.
Stigmas

included, lobes unequal, abaxial 1.5 times adaxial.

included, lobes equal.
2n

= 16.

= 16.

Diplacus graniticola

Diplacus vandenbergensis
Phenology Flowering Apr–Sep. Flowering Apr–Jun.
Habitat Granite cracks and crevices. Sandy open or disturbed areas, among shrubs.
Elevation 300–2100 m. (1000–6900 ft.) 70–150 m. (200–500 ft.)
Distribution
from FNA
CA
[WildflowerSearch map]
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
Discussion

Diplacus graniticola occurs in the Sierra Nevada from Tuolumne County to northern Tulare County. These plants previously were identified within D. layneae, with which they are partially sympatric; where these two occur together, D. layneae often grows in granite-derived sand and gravel immediately adjacent to the granite rock habitat of D. graniticola.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Diplacus vandenbergensis is endemic to Santa Barbara County, mostly on the north side of the city of Lompoc (La Purisima Mission State Historic Park; also on and near Vandenberg Air Force Base) plus one other locality about 10 kilometers farther west (Santa Ynez Valley, 8 miles west of Buellton). Thompson described Mimulus fremontii var. vandenbergensis as a geographically distinct variant of M. fremontii. The discontinuous morphological difference, allopatric/parapatric geographical distribution, and lack of intergrading populations support recognition of the plants with yellow corollas at specific rank. Plants with yellow corollas are found in D. bicolor, D. mephiticus, and D. parryi, but these morphs are found through most of the geographic ranges of these taxa and sometimes are intermixed in populations.

(Discussion copyrighted by Flora of North America; reprinted with permission.)
Source FNA vol. 17, p. 438. FNA vol. 17, p. 432.
Parent taxa Phrymaceae > Diplacus Phrymaceae > Diplacus
Sibling taxa
D. angustatus, D. aridus, D. aurantiacus, D. bicolor, D. bigelovii, D. bolanderi, D. brandegeei, D. brevipes, D. calycinus, D. cascadensis, D. clevelandii, D. clivicola, D. compactus, D. congdonii, D. constrictus, D. cusickii, D. cusickioides, D. deschutesensis, D. douglasii, D. fremontii, D. grandiflorus, D. jepsonii, D. johnstonii, D. kelloggii, D. layneae, D. leptaleus, D. linearis, D. longiflorus, D. mephiticus, D. mohavensis, D. nanus, D. ovatus, D. parryi, D. parviflorus, D. pictus, D. pulchellus, D. puniceus, D. pygmaeus, D. rattanii, D. rupicola, D. rutilus, D. thompsonii, D. torreyi, D. traskiae, D. tricolor, D. vandenbergensis, D. viscidus
D. angustatus, D. aridus, D. aurantiacus, D. bicolor, D. bigelovii, D. bolanderi, D. brandegeei, D. brevipes, D. calycinus, D. cascadensis, D. clevelandii, D. clivicola, D. compactus, D. congdonii, D. constrictus, D. cusickii, D. cusickioides, D. deschutesensis, D. douglasii, D. fremontii, D. grandiflorus, D. graniticola, D. jepsonii, D. johnstonii, D. kelloggii, D. layneae, D. leptaleus, D. linearis, D. longiflorus, D. mephiticus, D. mohavensis, D. nanus, D. ovatus, D. parryi, D. parviflorus, D. pictus, D. pulchellus, D. puniceus, D. pygmaeus, D. rattanii, D. rupicola, D. rutilus, D. thompsonii, D. torreyi, D. traskiae, D. tricolor, D. viscidus
Synonyms Mimulus fremontii var. vandenbergensis
Name authority Schoenig: Phytoneuron 2017-24: 1, figs. 1, 3–10. (2017) (D. M. Thompson) G. L. Nesom: Phytoneuron 2012-47: 2. (2012)
Web links 
Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).



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