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granite-crack monkeyflower

brownies, Douglas' monkeyflower, mouse-ear monkeyflower, purple mouse-ears

Habit Herbs, annual, herbage usually drying dark. Herbs, annual.
Stems

erect, 60–120(–150) mm, nodes 4–15(–20), internodes shorter than leaves, glandular-villous with gland-tipped hairs 1–1.6 mm.

erect, 3–40(–80) mm, glandular-puberulent and/or glandular-pubescent.

Leaves

usually cauline, relatively even-sized;

petiole weakly delimited;

blade usually lanceolate to ovate-lanceolate, 20–40 × 4–12 mm, margins entire, rarely toothed, plane, apex rounded to obtuse or acute, surfaces: proximals often glabrate abaxially, distals glandular-villous.

usually basal, reduced distally;

petiole absent, larger with petiole-like extension;

blade obovate to elliptic, 5–28(–35) × (2–)3–10(–12) mm, margins entire or crenate, plane, not ciliate, apex obtuse, surfaces: proximals glabrate, distals glandular-pilose adaxially.

Pedicels

1–3 mm in fruit.

(1–)2–4 mm in fruit, usually twisting to invert calyx.

Flowers

2 per node, or 1 or 2 per node on 1 plant, chasmogamous.

2 per node, or 1 or 2 per node on 1 plant, chasmogamous or cleistogamous.

Styles

glandular-puberulent.

pubescent distally.

Corollas

nearly white or pale lavender to pinkish or pale to dark magenta, each lobe with a dark medial line extending nearly to tip, throat with a dark red or purple splotch at junction of each abaxial lobe and adjacent lateral lobe, throat floor sometimes with 2 adjacent white splotches at lateral lobe bases, palate ridges yellow, tube-throat 15–20 mm, limb 10–16 mm diam., bilabiate.

throat magenta to dark purple, inside with longitudinal gold markings basally, adaxial lip magenta, palate ridges golden yellow, tube-throat (15–)20–41(–45) mm, limb 10–14 mm diam., abaxial lip essentially absent.

Calyces

symmetrically attached to pedicels, not inflated in fruit, 8–12 mm, glandular-villous, tube strongly plicate, lobes triangular, subequal, apex acute, ribs narrow, darkened, blackish, thickened, strongly raised, intercostal areas green to purple, not membranous.

distinctly asymmetrically attached to pedicel, not inflated in fruit, (6–)8–14(–15) mm, sparsely glandular-pilose, lobes subequal, apex appearing acute due to their continuity with ribs, ribs green to purplish, intercostal areas whitish.

Capsules

6–10 mm.

(2.5–)3–6.5 mm.

Anthers

included, ciliate.

(distal pair) nearly exserted, glabrous or glabrate.

Stigmas

included, lobes unequal, abaxial 1.5 times adaxial.

exserted, lobes unequal, abaxial 10–20 times adaxial.

2n

= 16.

= 18.

Diplacus graniticola

Diplacus douglasii

Phenology Flowering Apr–Sep. Flowering Feb–Apr(–May).
Habitat Granite cracks and crevices. Gentle to moderately steep slopes, upper banks of small creeks and ditches.
Elevation 300–2100 m. (1000–6900 ft.) 50–1200 m. (200–3900 ft.)
Distribution
from FNA
CA
[WildflowerSearch map]
from FNA
CA; OR
[WildflowerSearch map]
Discussion

Diplacus graniticola occurs in the Sierra Nevada from Tuolumne County to northern Tulare County. These plants previously were identified within D. layneae, with which they are partially sympatric; where these two occur together, D. layneae often grows in granite-derived sand and gravel immediately adjacent to the granite rock habitat of D. graniticola.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Diplacus douglasii occurs in southwestern Oregon and the northern two-thirds of California.

Diplacus douglasii is distinct in the complete or nearly complete lack of abaxial corolla lobes. Diplacus brandegeei, D. congdonii, D. kelloggii, and D. traskiae, the other species of sect. Cleisanthus, also show a distinct tendency toward reduction of the abaxial corolla lobes, and all species of the group except D. brandegeei produce markedly elongate corolla tubes.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 438. FNA vol. 17, p. 445.
Parent taxa Phrymaceae > Diplacus Phrymaceae > Diplacus
Sibling taxa
D. angustatus, D. aridus, D. aurantiacus, D. bicolor, D. bigelovii, D. bolanderi, D. brandegeei, D. brevipes, D. calycinus, D. cascadensis, D. clevelandii, D. clivicola, D. compactus, D. congdonii, D. constrictus, D. cusickii, D. cusickioides, D. deschutesensis, D. douglasii, D. fremontii, D. grandiflorus, D. jepsonii, D. johnstonii, D. kelloggii, D. layneae, D. leptaleus, D. linearis, D. longiflorus, D. mephiticus, D. mohavensis, D. nanus, D. ovatus, D. parryi, D. parviflorus, D. pictus, D. pulchellus, D. puniceus, D. pygmaeus, D. rattanii, D. rupicola, D. rutilus, D. thompsonii, D. torreyi, D. traskiae, D. tricolor, D. vandenbergensis, D. viscidus
D. angustatus, D. aridus, D. aurantiacus, D. bicolor, D. bigelovii, D. bolanderi, D. brandegeei, D. brevipes, D. calycinus, D. cascadensis, D. clevelandii, D. clivicola, D. compactus, D. congdonii, D. constrictus, D. cusickii, D. cusickioides, D. deschutesensis, D. fremontii, D. grandiflorus, D. graniticola, D. jepsonii, D. johnstonii, D. kelloggii, D. layneae, D. leptaleus, D. linearis, D. longiflorus, D. mephiticus, D. mohavensis, D. nanus, D. ovatus, D. parryi, D. parviflorus, D. pictus, D. pulchellus, D. puniceus, D. pygmaeus, D. rattanii, D. rupicola, D. rutilus, D. thompsonii, D. torreyi, D. traskiae, D. tricolor, D. vandenbergensis, D. viscidus
Synonyms Eunanus douglasii, Mimulus cleistogamus, M. douglasii
Name authority Schoenig: Phytoneuron 2017-24: 1, figs. 1, 3–10. (2017) (Bentham) G. L. Nesom: Phytoneuron 2012-39: 32. (2012)
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