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granite-crack monkeyflower

Cusick's monkeyflower, Nesom's monkey-flower

Habit Herbs, annual, herbage usually drying dark. Herbs, annual.
Stems

erect, 60–120(–150) mm, nodes 4–15(–20), internodes shorter than leaves, glandular-villous with gland-tipped hairs 1–1.6 mm.

erect to erect-ascending, (10–)30–240(–350) mm, distal internodes 1–5 mm, densely glandular-villous.

Leaves

usually cauline, relatively even-sized;

petiole weakly delimited;

blade usually lanceolate to ovate-lanceolate, 20–40 × 4–12 mm, margins entire, rarely toothed, plane, apex rounded to obtuse or acute, surfaces: proximals often glabrate abaxially, distals glandular-villous.

basal and cauline or usually cauline, relatively even-sized or gradually larger distally;

petiole absent, proximal base short petiole-like, 1–5 mm;

blade ovate to broadly elliptic-ovate, (10–)15–25(–35) × 4–16 mm, margins entire, plane, apex abruptly acuminate, surfaces densely glandular-villous.

Pedicels

1–3 mm in fruit.

1–1.5 mm in fruit.

Flowers

2 per node, or 1 or 2 per node on 1 plant, chasmogamous.

2 per node, or 1 or 2 per node on 1 plant, chasmogamous.

Styles

glandular-puberulent.

puberulent, at least on distal 1/2.

Corollas

nearly white or pale lavender to pinkish or pale to dark magenta, each lobe with a dark medial line extending nearly to tip, throat with a dark red or purple splotch at junction of each abaxial lobe and adjacent lateral lobe, throat floor sometimes with 2 adjacent white splotches at lateral lobe bases, palate ridges yellow, tube-throat 15–20 mm, limb 10–16 mm diam., bilabiate.

magenta or rose purple, tube yellow, throat usually yellow, throat and distal tube red-spotted on floor, palate ridges yellow, tube-throat 13–16(–19) mm, limb 14–26 mm diam., bilabiate.

Calyces

symmetrically attached to pedicels, not inflated in fruit, 8–12 mm, glandular-villous, tube strongly plicate, lobes triangular, subequal, apex acute, ribs narrow, darkened, blackish, thickened, strongly raised, intercostal areas green to purple, not membranous.

symmetrically attached to pedicels, not inflated in fruit, 7–12 mm, glabrous or minutely stipitate-glandular, lobes unequal, apex linear-acuminate, sharp-pointed, ribs green distally, intercostal areas whitish.

Capsules

6–10 mm.

10–17 mm.

Anthers

included, ciliate.

included, glabrous or sparsely hirsutulous.

Stigmas

included, lobes unequal, abaxial 1.5 times adaxial.

exserted, lobes subequal (herkogamous).

2n

= 16.

Diplacus graniticola

Diplacus cusickioides

Phenology Flowering Apr–Sep. Flowering May–Aug.
Habitat Granite cracks and crevices. Lava formations, steep slopes, roadsides, volcanic gravels, scree, ash.
Elevation 300–2100 m. (1000–6900 ft.) (400–)600–1500(–2000) m. ((1300–)2000–4900(–6600) ft.)
Distribution
from FNA
CA
[WildflowerSearch map]
from FNA
CA; ID; OR; WA
[WildflowerSearch map]
Discussion

Diplacus graniticola occurs in the Sierra Nevada from Tuolumne County to northern Tulare County. These plants previously were identified within D. layneae, with which they are partially sympatric; where these two occur together, D. layneae often grows in granite-derived sand and gravel immediately adjacent to the granite rock habitat of D. graniticola.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Diplacus cusickioides occurs in western Idaho, eastern Oregon, Klickitat County, Washington, and apparently in a disjunct population system in Modoc County, California, on the east side of the Warner Mountains.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 438. FNA vol. 17, p. 439.
Parent taxa Phrymaceae > Diplacus Phrymaceae > Diplacus
Sibling taxa
D. angustatus, D. aridus, D. aurantiacus, D. bicolor, D. bigelovii, D. bolanderi, D. brandegeei, D. brevipes, D. calycinus, D. cascadensis, D. clevelandii, D. clivicola, D. compactus, D. congdonii, D. constrictus, D. cusickii, D. cusickioides, D. deschutesensis, D. douglasii, D. fremontii, D. grandiflorus, D. jepsonii, D. johnstonii, D. kelloggii, D. layneae, D. leptaleus, D. linearis, D. longiflorus, D. mephiticus, D. mohavensis, D. nanus, D. ovatus, D. parryi, D. parviflorus, D. pictus, D. pulchellus, D. puniceus, D. pygmaeus, D. rattanii, D. rupicola, D. rutilus, D. thompsonii, D. torreyi, D. traskiae, D. tricolor, D. vandenbergensis, D. viscidus
D. angustatus, D. aridus, D. aurantiacus, D. bicolor, D. bigelovii, D. bolanderi, D. brandegeei, D. brevipes, D. calycinus, D. cascadensis, D. clevelandii, D. clivicola, D. compactus, D. congdonii, D. constrictus, D. cusickii, D. deschutesensis, D. douglasii, D. fremontii, D. grandiflorus, D. graniticola, D. jepsonii, D. johnstonii, D. kelloggii, D. layneae, D. leptaleus, D. linearis, D. longiflorus, D. mephiticus, D. mohavensis, D. nanus, D. ovatus, D. parryi, D. parviflorus, D. pictus, D. pulchellus, D. puniceus, D. pygmaeus, D. rattanii, D. rupicola, D. rutilus, D. thompsonii, D. torreyi, D. traskiae, D. tricolor, D. vandenbergensis, D. viscidus
Name authority Schoenig: Phytoneuron 2017-24: 1, figs. 1, 3–10. (2017) G. L. Nesom: Phytoneuron 2013-65: 6, figs. 3, 4. (2013)
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