FNA: Diplacus graniticola vs. Diplacus bolanderi

	Diplacus graniticola	Diplacus bolanderi
	granite-crack monkeyflower	Bolander's monkeyflower
Habit	Herbs, annual, herbage usually drying dark.	Herbs, annual.
Stems	erect, 60–120(–150) mm, nodes 4–15(–20), internodes shorter than leaves, glandular-villous with gland-tipped hairs 1–1.6 mm.	erect, 20–900(–1200) mm, usually glandular-pubescent and viscid.
Leaves	usually cauline, relatively even-sized; petiole weakly delimited; blade usually lanceolate to ovate-lanceolate, 20–40 × 4–12 mm, margins entire, rarely toothed, plane, apex rounded to obtuse or acute, surfaces: proximals often glabrate abaxially, distals glandular-villous.	basal and cauline, basal in rosette, cauline gradually reduced distally; petiole present proximally, usually absent distally; blade ovate, obovate, elliptic, or oblanceolate, 5–60(–85) × 1.5–25(–32) mm, margins entire or largest irregularly serrate, plane, apex rounded to obtuse, surfaces: proximals nearly glabrous, distals glandular-pubescent.
Pedicels	1–3 mm in fruit.	2–5(–8) mm in fruit.
Flowers	2 per node, or 1 or 2 per node on 1 plant, chasmogamous.	2 per node, or 1 or 2 per node on 1 plant (1 per node on depauperate plants), chasmogamous.
Styles	glandular-puberulent.	glandular-puberulent.
Corollas	nearly white or pale lavender to pinkish or pale to dark magenta, each lobe with a dark medial line extending nearly to tip, throat with a dark red or purple splotch at junction of each abaxial lobe and adjacent lateral lobe, throat floor sometimes with 2 adjacent white splotches at lateral lobe bases, palate ridges yellow, tube-throat 15–20 mm, limb 10–16 mm diam., bilabiate.	magenta, throat often darkening near floor, without dark spots, palate ridges or floor white, usually magenta-speckled, tube-throat 10–30 mm, limb 6–20 mm diam., not bilabiate.
Calyces	symmetrically attached to pedicels, not inflated in fruit, 8–12 mm, glandular-villous, tube strongly plicate, lobes triangular, subequal, apex acute, ribs narrow, darkened, blackish, thickened, strongly raised, intercostal areas green to purple, not membranous.	symmetrically attached to pedicels, inflated in fruit, (7–)9–25(–27) mm, glandular-pubescent and viscid, lobes often spreading in fruit, unequal if calyx large, apex acute to attenuate, ribs prominent, purplish, intercostal areas white.
Capsules	6–10 mm.	8–20 mm.
Anthers	included, ciliate.	included, glabrous.
Stigmas	included, lobes unequal, abaxial 1.5 times adaxial.	exserted, lobes unequal, abaxial 2–3 times adaxial.
2n	= 16.	= 16.

	Diplacus graniticola	Diplacus bolanderi
Phenology	Flowering Apr–Sep.	Flowering Apr–Jul.
Habitat	Granite cracks and crevices.	Granite outcrops, slopes with decomposed granite soils.
Elevation	300–2100 m. (1000–6900 ft.)	(100–)300–1700(–2000) m. ((300–)1000–5600(–6600) ft.)
Distribution	CA [WildflowerSearch map]	CA; OR [WildflowerSearch map]
Discussion	Diplacus graniticola occurs in the Sierra Nevada from Tuolumne County to northern Tulare County. These plants previously were identified within D. layneae, with which they are partially sympatric; where these two occur together, D. layneae often grows in granite-derived sand and gravel immediately adjacent to the granite rock habitat of D. graniticola. (Discussion copyrighted by Flora of North America; reprinted with permission.)	Diplacus bolanderi is scattered and uncommon through most of California (with concentrations in the southern North Coast Ranges and Central Sierra) and apparently is disjunct in Jackson County, Oregon. Diplacus bolanderi is highly variable in height, leaf size, corolla size, and calyx size. Identity is unmistakable in larger plants; the inflated calyx with lobes strongly unequal in length usually is helpful in identification of smaller plants. (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 17, p. 438.	FNA vol. 17, p. 434.
Parent taxa	Phrymaceae > Diplacus	Phrymaceae > Diplacus
Sibling taxa	D. angustatus, D. aridus, D. aurantiacus, D. bicolor, D. bigelovii, D. bolanderi, D. brandegeei, D. brevipes, D. calycinus, D. cascadensis, D. clevelandii, D. clivicola, D. compactus, D. congdonii, D. constrictus, D. cusickii, D. cusickioides, D. deschutesensis, D. douglasii, D. fremontii, D. grandiflorus, D. jepsonii, D. johnstonii, D. kelloggii, D. layneae, D. leptaleus, D. linearis, D. longiflorus, D. mephiticus, D. mohavensis, D. nanus, D. ovatus, D. parryi, D. parviflorus, D. pictus, D. pulchellus, D. puniceus, D. pygmaeus, D. rattanii, D. rupicola, D. rutilus, D. thompsonii, D. torreyi, D. traskiae, D. tricolor, D. vandenbergensis, D. viscidus	D. angustatus, D. aridus, D. aurantiacus, D. bicolor, D. bigelovii, D. brandegeei, D. brevipes, D. calycinus, D. cascadensis, D. clevelandii, D. clivicola, D. compactus, D. congdonii, D. constrictus, D. cusickii, D. cusickioides, D. deschutesensis, D. douglasii, D. fremontii, D. grandiflorus, D. graniticola, D. jepsonii, D. johnstonii, D. kelloggii, D. layneae, D. leptaleus, D. linearis, D. longiflorus, D. mephiticus, D. mohavensis, D. nanus, D. ovatus, D. parryi, D. parviflorus, D. pictus, D. pulchellus, D. puniceus, D. pygmaeus, D. rattanii, D. rupicola, D. rutilus, D. thompsonii, D. torreyi, D. traskiae, D. tricolor, D. vandenbergensis, D. viscidus
Synonyms		Mimulus bolanderi, M. platylaemus
Name authority	Schoenig: Phytoneuron 2017-24: 1, figs. 1, 3–10. (2017)	(A. Gray) G. L. Nesom: Phytoneuron 2012-39: 28. (2012)
Web links	Local floras: CA Local Web sites: CalFlora, CalPhotos WildflowerSearch iNaturalist (observations) LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	Local floras: CA, OR Local Web sites: CalFlora, CalPhotos, Flora NW, PNW Herbaria WildflowerSearch iNaturalist (observations) LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Diplacus bolanderi

granite-crack monkeyflower

Bolander's monkeyflower

Habit

Herbs, annual, herbage usually drying dark.

Herbs, annual.

Stems

erect, 60–120(–150) mm, nodes 4–15(–20), internodes shorter than leaves, glandular-villous with gland-tipped hairs 1–1.6 mm.

erect, 20–900(–1200) mm, usually glandular-pubescent and viscid.

Leaves

usually cauline, relatively even-sized;

petiole weakly delimited;

blade usually lanceolate to ovate-lanceolate, 20–40 × 4–12 mm, margins entire, rarely toothed, plane, apex rounded to obtuse or acute, surfaces: proximals often glabrate abaxially, distals glandular-villous.

basal and cauline, basal in rosette, cauline gradually reduced distally;

petiole present proximally, usually absent distally;

blade ovate, obovate, elliptic, or oblanceolate, 5–60(–85) × 1.5–25(–32) mm, margins entire or largest irregularly serrate, plane, apex rounded to obtuse, surfaces: proximals nearly glabrous, distals glandular-pubescent.

Pedicels

1–3 mm in fruit.

2–5(–8) mm in fruit.

Flowers

2 per node, or 1 or 2 per node on 1 plant, chasmogamous.

2 per node, or 1 or 2 per node on 1 plant (1 per node on depauperate plants), chasmogamous.

Styles

glandular-puberulent.

Corollas

nearly white or pale lavender to pinkish or pale to dark magenta, each lobe with a dark medial line extending nearly to tip, throat with a dark red or purple splotch at junction of each abaxial lobe and adjacent lateral lobe, throat floor sometimes with 2 adjacent white splotches at lateral lobe bases, palate ridges yellow, tube-throat 15–20 mm, limb 10–16 mm diam., bilabiate.

magenta, throat often darkening near floor, without dark spots, palate ridges or floor white, usually magenta-speckled, tube-throat 10–30 mm, limb 6–20 mm diam., not bilabiate.

Calyces

symmetrically attached to pedicels, not inflated in fruit, 8–12 mm, glandular-villous, tube strongly plicate, lobes triangular, subequal, apex acute, ribs narrow, darkened, blackish, thickened, strongly raised, intercostal areas green to purple, not membranous.

symmetrically attached to pedicels, inflated in fruit, (7–)9–25(–27) mm, glandular-pubescent and viscid, lobes often spreading in fruit, unequal if calyx large, apex acute to attenuate, ribs prominent, purplish, intercostal areas white.

Capsules

6–10 mm.

8–20 mm.

Anthers

included, ciliate.

included, glabrous.

Stigmas

included, lobes unequal, abaxial 1.5 times adaxial.

exserted, lobes unequal, abaxial 2–3 times adaxial.

= 16.

Diplacus graniticola

Diplacus bolanderi

Phenology

Flowering Apr–Sep.

Flowering Apr–Jul.

Habitat

Granite cracks and crevices.

Granite outcrops, slopes with decomposed granite soils.

Elevation

300–2100 m. (1000–6900 ft.)

(100–)300–1700(–2000) m. ((300–)1000–5600(–6600) ft.)

Distribution

[WildflowerSearch map]

CA; OR

[WildflowerSearch map]

Discussion

Diplacus graniticola occurs in the Sierra Nevada from Tuolumne County to northern Tulare County. These plants previously were identified within D. layneae, with which they are partially sympatric; where these two occur together, D. layneae often grows in granite-derived sand and gravel immediately adjacent to the granite rock habitat of D. graniticola.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Diplacus bolanderi is scattered and uncommon through most of California (with concentrations in the southern North Coast Ranges and Central Sierra) and apparently is disjunct in Jackson County, Oregon.

Diplacus bolanderi is highly variable in height, leaf size, corolla size, and calyx size. Identity is unmistakable in larger plants; the inflated calyx with lobes strongly unequal in length usually is helpful in identification of smaller plants.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 17, p. 438.

FNA vol. 17, p. 434.

Parent taxa

Phrymaceae > Diplacus

Sibling taxa

D. angustatus, D. aridus, D. aurantiacus, D. bicolor, D. bigelovii, D. bolanderi, D. brandegeei, D. brevipes, D. calycinus, D. cascadensis, D. clevelandii, D. clivicola, D. compactus, D. congdonii, D. constrictus, D. cusickii, D. cusickioides, D. deschutesensis, D. douglasii, D. fremontii, D. grandiflorus, D. jepsonii, D. johnstonii, D. kelloggii, D. layneae, D. leptaleus, D. linearis, D. longiflorus, D. mephiticus, D. mohavensis, D. nanus, D. ovatus, D. parryi, D. parviflorus, D. pictus, D. pulchellus, D. puniceus, D. pygmaeus, D. rattanii, D. rupicola, D. rutilus, D. thompsonii, D. torreyi, D. traskiae, D. tricolor, D. vandenbergensis, D. viscidus

D. angustatus, D. aridus, D. aurantiacus, D. bicolor, D. bigelovii, D. brandegeei, D. brevipes, D. calycinus, D. cascadensis, D. clevelandii, D. clivicola, D. compactus, D. congdonii, D. constrictus, D. cusickii, D. cusickioides, D. deschutesensis, D. douglasii, D. fremontii, D. grandiflorus, D. graniticola, D. jepsonii, D. johnstonii, D. kelloggii, D. layneae, D. leptaleus, D. linearis, D. longiflorus, D. mephiticus, D. mohavensis, D. nanus, D. ovatus, D. parryi, D. parviflorus, D. pictus, D. pulchellus, D. puniceus, D. pygmaeus, D. rattanii, D. rupicola, D. rutilus, D. thompsonii, D. torreyi, D. traskiae, D. tricolor, D. vandenbergensis, D. viscidus

Synonyms

Mimulus bolanderi, M. platylaemus

Name authority

Schoenig: Phytoneuron 2017-24: 1, figs. 1, 3–10. (2017)

(A. Gray) G. L. Nesom: Phytoneuron 2012-39: 28. (2012)

Web links

Local floras: CA, OR
Local Web sites: CalFlora, CalPhotos, Flora NW, PNW Herbaria
WildflowerSearch
iNaturalist (observations)
LBJ Wildflower Center
SEINet
Plants of the World Online
Encyclopedia of Life
Wikipedia
Google Image Search

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

Choose a data source: Flora of North America, OregonFlora

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Flora of North America species comparison

Diplacus graniticola

Diplacus bolanderi

Diplacus graniticola

Diplacus bolanderi