Diplacus deschutesensis |
Diplacus rutilus |
|
---|---|---|
Deschutes monkeyflower |
Santa Susana bush monkeyflower |
|
Habit | Herbs, annual. | Subshrubs. |
Stems | erect to erect-ascending, 40–150 mm, distal internodes 1–2 mm, minutely glandular-puberulent. |
erect, 300–1000(–2000) mm, glandular-puberulent and short-villous. |
Leaves | usually cauline, relatively even-sized or gradually larger distally; petiole absent, proximal base short petiole-like; blade broadly ovate or obovate to elliptic-ovate or elliptic-oblanceolate, 10–15(–25) × 4–13 mm, margins entire, plane, apex acuminate or cuspidate, surfaces glabrous or sparsely glandular-puberulent. |
usually cauline, relatively even-sized; petiole absent; blade elliptic to lanceolate, elliptic-lanceolate, or elliptic-oblanceolate, 25–65(–80) × 4–15(–25) mm, margins entire or serrate, revolute, apex acute to obtuse, abaxial surfaces densely hairy, hairs branched, adaxial glabrescent. |
Pedicels | 1–1.5 mm in fruit. |
5–16 mm in fruit. |
Flowers | usually from proximalmost to distal nodes, 2 per node, or 1 or 2 per node on 1 plant, chasmogamous. |
2 per node, chasmogamous. |
Styles | puberulent, at least on distal 1/2. |
minutely glandular. |
Corollas | light pink to magenta or rose purple, usually with a darker narrow line extending from throat onto each lobe midvein, throat yellow, palate ridges yellow, tube-throat 8–12 mm, limb 10–16 mm diam., bilabiate. |
red to orange-red, throat whitish at least on floor, palate ridges orange or white with orange crest, tube-throat 34–45 mm, limb (25–)28–40 mm diam., bilabiate, lobes oblong, apex of adaxial 2 each shallowly, asymmetrically incised. |
Calyces | symmetrically attached to pedicels, not inflated in fruit, 7–8 mm, glandular-puberulent, lobes subequal, apex linear-acuminate, ribs green distally, intercostal areas whitish. |
not inflated in fruit, 22–32 mm, glandular-puberulent and short glandular-villous to hirsute-villous, tube slightly dilated distally, lobes unequal, apex acute, ribs green, intercostal areas light green. |
Capsules | 7–9 mm. |
18–28 mm. |
Anthers | included, glabrous or sparsely hispidulous. |
included, glabrous. |
Stigmas | exserted, lobes subequal, abaxial slightly longer. |
included, lobes equal. |
2n | = 16. |
|
Diplacus deschutesensis |
Diplacus rutilus |
|
Phenology | Flowering Jun–Aug. | Flowering Mar–Jun. |
Habitat | Sandy and ashy soils, pumice sand and gravel, red clay slopes, hillsides, roadsides, bare areas, sagebrush, sagebrush-juniper, juniper, yellow pine and lodgepole pine forests. | Chaparral. |
Elevation | 700–1500 m. (2300–4900 ft.) | 400–600 m. (1300–2000 ft.) |
Distribution |
OR
|
CA |
Discussion | Diplacus deschutesensis is endemic to Crook, Deschutes, Jefferson, Klamath, Lake, and Wheeler counties of central Oregon. D. M. Thompson (2005) regarded these plants as a zone of stabilized hybrids, intermediate between Mimulus cusickii and typical M. nanus, the range just outside and west of the wider range of typical M. cusickii. In an earlier study that included both of the latter species, W. L. Ezell (1971, and by annotation in 1987) identified the same set of plants simply as M. cusickii, not associating them at all with M. nanus. A. L. Grant (1924, and by annotation of MO collections) identified them variously as either M. cusickii or M. ovatus. Thompson did not say what features of intermediacy he observed in the putative hybrids, but he did note that they produced leaves with acuminate-cuspidate apices and that they would key to M. cusickii. Leaves of Diplacus deschutesensis are broad with abruptly and sharply acuminate apices like those of D. cusickii, and the corolla coloration also is similar. The flowers (calyx length, corolla tube-throat length, limb width) and capsules of D. deschutesensis are considerably smaller, and the distal leaves are smaller with glabrous surfaces. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
H. E. McMinn (1951), M. C. Tulig and G. L Nesom (2012), and Nesom (2013c) treated Diplacus rutilus at specific rank, but P. A. Munz and D. D. Keck (1973) followed the original assessment of A. L. Grant (1924) in treating it as a variety of the light orange- to pale yellow-orange-flowered D. longiflorus, while R. M. Beeks (1962) and D. M. Thompson (2005) regarded this taxon as only a variant of D. longiflorus, without formal rank. It is maintained here as a distinct, red-flowered species localized in Los Angeles County, California. Diplacus rutilus occurs in pockets in a strip from Whittier and Pomona through North Pasadena westward to near the Ventura County line (the Santa Susana area being the type locality), a distance of almost 60 miles. Very few of these red-flowered collections can be separated in any feature except corolla color from typical Diplacus longiflorus. Red corollas have not been observed in D. longiflorus outside of Los Angeles County, and D. rutilus might be interpreted as reflecting local introgression in this area from D. puniceus, but the distinctive and tightly coherent geographical distribution of these red-flowered plants and their apparent absence elsewhere in the area where D. ×australis occurs suggest that the origin of D. rutilus is different from that of the highly variable D. ×australis. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 440. | FNA vol. 17, p. 451. |
Parent taxa | Phrymaceae > Diplacus | Phrymaceae > Diplacus |
Sibling taxa | ||
Synonyms | Mimulus longiflorus var. rutilus, D. longiflorus var. rutilus | |
Name authority | G. L. Nesom: Phytoneuron 2013-65: 8, fig. 5. (2013) | (A. L. Grant) McMinn: Madroño 11: 83. (1951) |
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