FNA: Diplacus deschutesensis vs. Diplacus clevelandii

	Diplacus deschutesensis	Diplacus clevelandii
	Deschutes monkeyflower	Cleveland's bush monkeyflower
Habit	Herbs, annual.	Herbs, perennial, sometimes rhizomatous.
Stems	erect to erect-ascending, 40–150 mm, distal internodes 1–2 mm, minutely glandular-puberulent.	erect, (200–)300–950(–1250) mm, glandular-villous.
Leaves	usually cauline, relatively even-sized or gradually larger distally; petiole absent, proximal base short petiole-like; blade broadly ovate or obovate to elliptic-ovate or elliptic-oblanceolate, 10–15(–25) × 4–13 mm, margins entire, plane, apex acuminate or cuspidate, surfaces glabrous or sparsely glandular-puberulent.	basal and cauline, relatively even-sized; petiole absent; blade lanceolate to oblong-lanceolate or elliptic-lanceolate, 20–110(–130) × 5–33(–40) mm, margins dentate to serrate, plane or revolute, apex acute, surfaces finely pubescent-glandular.
Pedicels	1–1.5 mm in fruit.	2–4(–7) mm in fruit.
Flowers	usually from proximalmost to distal nodes, 2 per node, or 1 or 2 per node on 1 plant, chasmogamous.	2 per node, chasmogamous.
Styles	puberulent, at least on distal 1/2.	minutely glandular.
Corollas	light pink to magenta or rose purple, usually with a darker narrow line extending from throat onto each lobe midvein, throat yellow, palate ridges yellow, tube-throat 8–12 mm, limb 10–16 mm diam., bilabiate.	bright golden yellow, often with reddish brown speckling on throat floor, palate ridges yellow, tube-throat 21–40 mm, limb 25–30 mm diam., nearly regular, lobes obovate-oblong.
Calyces	symmetrically attached to pedicels, not inflated in fruit, 7–8 mm, glandular-puberulent, lobes subequal, apex linear-acuminate, ribs green distally, intercostal areas whitish.	not inflated in fruit, 20–35(–37) mm, densely glandular-pubescent, tube slightly dilated distally, lobes subequal, apex acute.
Capsules	7–9 mm.	8–15(–17) mm.
Anthers	included, glabrous or sparsely hispidulous.	included, glabrous.
Stigmas	exserted, lobes subequal, abaxial slightly longer.	included, lobes unequal, abaxial 2 times longer than adaxial.
2n	= 16.	= 20.

	Diplacus deschutesensis	Diplacus clevelandii
Phenology	Flowering Jun–Aug.	Flowering Apr–Jun(–Jul).
Habitat	Sandy and ashy soils, pumice sand and gravel, red clay slopes, hillsides, roadsides, bare areas, sagebrush, sagebrush-juniper, juniper, yellow pine and lodgepole pine forests.	Dry, rocky openings in chaparral, roadcuts.
Elevation	700–1500 m. (2300–4900 ft.)	900–1500 m. (3000–4900 ft.)
Distribution	OR [WildflowerSearch map]	CA; Mexico (Baja California) [WildflowerSearch map] [BONAP county map]
Discussion	Diplacus deschutesensis is endemic to Crook, Deschutes, Jefferson, Klamath, Lake, and Wheeler counties of central Oregon. D. M. Thompson (2005) regarded these plants as a zone of stabilized hybrids, intermediate between Mimulus cusickii and typical M. nanus, the range just outside and west of the wider range of typical M. cusickii. In an earlier study that included both of the latter species, W. L. Ezell (1971, and by annotation in 1987) identified the same set of plants simply as M. cusickii, not associating them at all with M. nanus. A. L. Grant (1924, and by annotation of MO collections) identified them variously as either M. cusickii or M. ovatus. Thompson did not say what features of intermediacy he observed in the putative hybrids, but he did note that they produced leaves with acuminate-cuspidate apices and that they would key to M. cusickii. Leaves of Diplacus deschutesensis are broad with abruptly and sharply acuminate apices like those of D. cusickii, and the corolla coloration also is similar. The flowers (calyx length, corolla tube-throat length, limb width) and capsules of D. deschutesensis are considerably smaller, and the distal leaves are smaller with glabrous surfaces. (Discussion copyrighted by Flora of North America; reprinted with permission.)	Diplacus clevelandii is restricted to Orange, Riverside, and San Diego counties and in adjacent Mexico. (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 17, p. 440.	FNA vol. 17, p. 448.
Parent taxa	Phrymaceae > Diplacus	Phrymaceae > Diplacus
Sibling taxa	D. angustatus, D. aridus, D. aurantiacus, D. bicolor, D. bigelovii, D. bolanderi, D. brandegeei, D. brevipes, D. calycinus, D. cascadensis, D. clevelandii, D. clivicola, D. compactus, D. congdonii, D. constrictus, D. cusickii, D. cusickioides, D. douglasii, D. fremontii, D. grandiflorus, D. graniticola, D. jepsonii, D. johnstonii, D. kelloggii, D. layneae, D. leptaleus, D. linearis, D. longiflorus, D. mephiticus, D. mohavensis, D. nanus, D. ovatus, D. parryi, D. parviflorus, D. pictus, D. pulchellus, D. puniceus, D. pygmaeus, D. rattanii, D. rupicola, D. rutilus, D. thompsonii, D. torreyi, D. traskiae, D. tricolor, D. vandenbergensis, D. viscidus	D. angustatus, D. aridus, D. aurantiacus, D. bicolor, D. bigelovii, D. bolanderi, D. brandegeei, D. brevipes, D. calycinus, D. cascadensis, D. clivicola, D. compactus, D. congdonii, D. constrictus, D. cusickii, D. cusickioides, D. deschutesensis, D. douglasii, D. fremontii, D. grandiflorus, D. graniticola, D. jepsonii, D. johnstonii, D. kelloggii, D. layneae, D. leptaleus, D. linearis, D. longiflorus, D. mephiticus, D. mohavensis, D. nanus, D. ovatus, D. parryi, D. parviflorus, D. pictus, D. pulchellus, D. puniceus, D. pygmaeus, D. rattanii, D. rupicola, D. rutilus, D. thompsonii, D. torreyi, D. traskiae, D. tricolor, D. vandenbergensis, D. viscidus
Synonyms		Mimulus clevelandii
Name authority	G. L. Nesom: Phytoneuron 2013-65: 8, fig. 5. (2013)	(Brandegee) Greene: Erythea 4: 22. (1896) — (as clevelandi)
Web links	Local floras: OR Local Web sites: Flora NW, PNW Herbaria WildflowerSearch iNaturalist (observations) LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	Local floras: CA Local Web sites: CalFlora, CalPhotos WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Diplacus clevelandii

Deschutes monkeyflower

Cleveland's bush monkeyflower

Habit

Herbs, annual.

Herbs, perennial, sometimes rhizomatous.

Stems

erect to erect-ascending, 40–150 mm, distal internodes 1–2 mm, minutely glandular-puberulent.

erect, (200–)300–950(–1250) mm, glandular-villous.

Leaves

usually cauline, relatively even-sized or gradually larger distally;

petiole absent, proximal base short petiole-like;

blade broadly ovate or obovate to elliptic-ovate or elliptic-oblanceolate, 10–15(–25) × 4–13 mm, margins entire, plane, apex acuminate or cuspidate, surfaces glabrous or sparsely glandular-puberulent.

basal and cauline, relatively even-sized;

petiole absent;

blade lanceolate to oblong-lanceolate or elliptic-lanceolate, 20–110(–130) × 5–33(–40) mm, margins dentate to serrate, plane or revolute, apex acute, surfaces finely pubescent-glandular.

Pedicels

1–1.5 mm in fruit.

2–4(–7) mm in fruit.

Flowers

usually from proximalmost to distal nodes, 2 per node, or 1 or 2 per node on 1 plant, chasmogamous.

2 per node, chasmogamous.

Styles

puberulent, at least on distal 1/2.

minutely glandular.

Corollas

light pink to magenta or rose purple, usually with a darker narrow line extending from throat onto each lobe midvein, throat yellow, palate ridges yellow, tube-throat 8–12 mm, limb 10–16 mm diam., bilabiate.

bright golden yellow, often with reddish brown speckling on throat floor, palate ridges yellow, tube-throat 21–40 mm, limb 25–30 mm diam., nearly regular, lobes obovate-oblong.

Calyces

symmetrically attached to pedicels, not inflated in fruit, 7–8 mm, glandular-puberulent, lobes subequal, apex linear-acuminate, ribs green distally, intercostal areas whitish.

not inflated in fruit, 20–35(–37) mm, densely glandular-pubescent, tube slightly dilated distally, lobes subequal, apex acute.

Capsules

7–9 mm.

8–15(–17) mm.

Anthers

included, glabrous or sparsely hispidulous.

included, glabrous.

Stigmas

exserted, lobes subequal, abaxial slightly longer.

included, lobes unequal, abaxial 2 times longer than adaxial.

= 16.

= 20.

Diplacus deschutesensis

Diplacus clevelandii

Phenology

Flowering Jun–Aug.

Flowering Apr–Jun(–Jul).

Habitat

Sandy and ashy soils, pumice sand and gravel, red clay slopes, hillsides, roadsides, bare areas, sagebrush, sagebrush-juniper, juniper, yellow pine and lodgepole pine forests.

Dry, rocky openings in chaparral, roadcuts.

Elevation

700–1500 m. (2300–4900 ft.)

900–1500 m. (3000–4900 ft.)

Distribution

[WildflowerSearch map]

CA; Mexico (Baja California)

[WildflowerSearch map]

[BONAP county map]

Discussion

Diplacus deschutesensis is endemic to Crook, Deschutes, Jefferson, Klamath, Lake, and Wheeler counties of central Oregon. D. M. Thompson (2005) regarded these plants as a zone of stabilized hybrids, intermediate between Mimulus cusickii and typical M. nanus, the range just outside and west of the wider range of typical M. cusickii. In an earlier study that included both of the latter species, W. L. Ezell (1971, and by annotation in 1987) identified the same set of plants simply as M. cusickii, not associating them at all with M. nanus. A. L. Grant (1924, and by annotation of MO collections) identified them variously as either M. cusickii or M. ovatus. Thompson did not say what features of intermediacy he observed in the putative hybrids, but he did note that they produced leaves with acuminate-cuspidate apices and that they would key to M. cusickii.

Leaves of Diplacus deschutesensis are broad with abruptly and sharply acuminate apices like those of D. cusickii, and the corolla coloration also is similar. The flowers (calyx length, corolla tube-throat length, limb width) and capsules of D. deschutesensis are considerably smaller, and the distal leaves are smaller with glabrous surfaces.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Diplacus clevelandii is restricted to Orange, Riverside, and San Diego counties and in adjacent Mexico.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 17, p. 440.

FNA vol. 17, p. 448.

Parent taxa

Phrymaceae > Diplacus

Sibling taxa

D. angustatus, D. aridus, D. aurantiacus, D. bicolor, D. bigelovii, D. bolanderi, D. brandegeei, D. brevipes, D. calycinus, D. cascadensis, D. clevelandii, D. clivicola, D. compactus, D. congdonii, D. constrictus, D. cusickii, D. cusickioides, D. douglasii, D. fremontii, D. grandiflorus, D. graniticola, D. jepsonii, D. johnstonii, D. kelloggii, D. layneae, D. leptaleus, D. linearis, D. longiflorus, D. mephiticus, D. mohavensis, D. nanus, D. ovatus, D. parryi, D. parviflorus, D. pictus, D. pulchellus, D. puniceus, D. pygmaeus, D. rattanii, D. rupicola, D. rutilus, D. thompsonii, D. torreyi, D. traskiae, D. tricolor, D. vandenbergensis, D. viscidus

D. angustatus, D. aridus, D. aurantiacus, D. bicolor, D. bigelovii, D. bolanderi, D. brandegeei, D. brevipes, D. calycinus, D. cascadensis, D. clivicola, D. compactus, D. congdonii, D. constrictus, D. cusickii, D. cusickioides, D. deschutesensis, D. douglasii, D. fremontii, D. grandiflorus, D. graniticola, D. jepsonii, D. johnstonii, D. kelloggii, D. layneae, D. leptaleus, D. linearis, D. longiflorus, D. mephiticus, D. mohavensis, D. nanus, D. ovatus, D. parryi, D. parviflorus, D. pictus, D. pulchellus, D. puniceus, D. pygmaeus, D. rattanii, D. rupicola, D. rutilus, D. thompsonii, D. torreyi, D. traskiae, D. tricolor, D. vandenbergensis, D. viscidus

Synonyms

Mimulus clevelandii

Name authority

G. L. Nesom: Phytoneuron 2013-65: 8, fig. 5. (2013)

(Brandegee) Greene: Erythea 4: 22. (1896) — (as clevelandi)

Web links

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

Choose a data source: Flora of North America, OregonFlora

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Flora of North America species comparison

Diplacus deschutesensis

Diplacus clevelandii

Diplacus deschutesensis

Diplacus clevelandii