Diplacus deschutesensis |
Diplacus brandegeei |
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Deschutes monkeyflower |
Santa Cruz Island monkeyflower, Santa Cruz Island or broadleaf monkeyflower |
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Habit | Herbs, annual. | Herbs, annual. |
Stems | erect to erect-ascending, 40–150 mm, distal internodes 1–2 mm, minutely glandular-puberulent. |
erect, 10–100 mm, glandular-puberulent to glandular-pubescent. |
Leaves | usually cauline, relatively even-sized or gradually larger distally; petiole absent, proximal base short petiole-like; blade broadly ovate or obovate to elliptic-ovate or elliptic-oblanceolate, 10–15(–25) × 4–13 mm, margins entire, plane, apex acuminate or cuspidate, surfaces glabrous or sparsely glandular-puberulent. |
mostly cauline, relatively even-sized; petiole absent, larger with petiole-like extension; blade narrowly to broadly ovate to broadly elliptic, (4–)8–39 × (2–)4–23 mm, margins entire or crenate, plane, proximal 1/2 usually ciliate, apex obtuse, surfaces glandular-pubescent. |
Pedicels | 1–1.5 mm in fruit. |
1.5–4 mm in fruit, not twisting to invert calyx. |
Flowers | usually from proximalmost to distal nodes, 2 per node, or 1 or 2 per node on 1 plant, chasmogamous. |
2 per node, or 1 or 2 per node on 1 plant, chasmogamous. |
Styles | puberulent, at least on distal 1/2. |
pubescent distally. |
Corollas | light pink to magenta or rose purple, usually with a darker narrow line extending from throat onto each lobe midvein, throat yellow, palate ridges yellow, tube-throat 8–12 mm, limb 10–16 mm diam., bilabiate. |
throat magenta, golden yellow at base, limb magenta, palate ridges golden yellow, purple-speckled, tube-throat 12–19 mm, limb 5–11 mm diam., bilabiate, abaxial lip smaller than adaxial. |
Calyces | symmetrically attached to pedicels, not inflated in fruit, 7–8 mm, glandular-puberulent, lobes subequal, apex linear-acuminate, ribs green distally, intercostal areas whitish. |
distinctly asymmetrically attached to pedicel, inflated in fruit, 8–13 mm, glandular-pubescent, lobes subequal, apex obtuse, ribs green, intercostal areas pale green to whitish. |
Capsules | 7–9 mm. |
(5–)6–11 mm, indehiscent. |
Anthers | included, glabrous or sparsely hispidulous. |
(distal pair) nearly exserted, glabrous. |
Stigmas | exserted, lobes subequal, abaxial slightly longer. |
exserted, lobes unequal, abaxial 3–4 times adaxial. |
2n | = 16. |
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Diplacus deschutesensis |
Diplacus brandegeei |
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Phenology | Flowering Jun–Aug. | Flowering Mar–Apr(–May). |
Habitat | Sandy and ashy soils, pumice sand and gravel, red clay slopes, hillsides, roadsides, bare areas, sagebrush, sagebrush-juniper, juniper, yellow pine and lodgepole pine forests. | Rocky, brushy slopes. |
Elevation | 700–1500 m. (2300–4900 ft.) | 100–300 m. (300–1000 ft.) |
Distribution |
OR
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CA; Mexico (Baja California) |
Discussion | Diplacus deschutesensis is endemic to Crook, Deschutes, Jefferson, Klamath, Lake, and Wheeler counties of central Oregon. D. M. Thompson (2005) regarded these plants as a zone of stabilized hybrids, intermediate between Mimulus cusickii and typical M. nanus, the range just outside and west of the wider range of typical M. cusickii. In an earlier study that included both of the latter species, W. L. Ezell (1971, and by annotation in 1987) identified the same set of plants simply as M. cusickii, not associating them at all with M. nanus. A. L. Grant (1924, and by annotation of MO collections) identified them variously as either M. cusickii or M. ovatus. Thompson did not say what features of intermediacy he observed in the putative hybrids, but he did note that they produced leaves with acuminate-cuspidate apices and that they would key to M. cusickii. Leaves of Diplacus deschutesensis are broad with abruptly and sharply acuminate apices like those of D. cusickii, and the corolla coloration also is similar. The flowers (calyx length, corolla tube-throat length, limb width) and capsules of D. deschutesensis are considerably smaller, and the distal leaves are smaller with glabrous surfaces. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Diplacus brandegeei is known only from Santa Catalina (Los Angeles County), Santa Cruz (Santa Barbara County), and Guadalupe (Baja California) islands. The earliest name for the species, Mimulus latifolius A. Gray, was first transferred to Diplacus as D. latifolius (A. Gray) G. L. Nesom, but that name is a later homonym of D. latifolius Nuttall. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 440. | FNA vol. 17, p. 446. |
Parent taxa | Phrymaceae > Diplacus | Phrymaceae > Diplacus |
Sibling taxa | ||
Synonyms | Mimulus brandegeei, M. latifolius | |
Name authority | G. L. Nesom: Phytoneuron 2013-65: 8, fig. 5. (2013) | (Pennell) G. L. Nesom: Phytoneuron 2012-54: 1. (2012) |
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