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Cusick's monkey-flower

sticky monkeyflower, viscid monkeyflower

Habit Herbs, annual. Herbs, annual.
Stems

erect to erect-ascending, 10–80 mm, distal internodes 2–20 mm, minutely glandular-puberulent.

erect, (30–)60–370 mm, densely glandular-pubescent with viscid hairs.

Leaves

basal and cauline or usually cauline, relatively even-sized or gradually larger distally;

petiole absent, proximal base short petiole-like, 1–5 mm;

blade ovate to broadly elliptic-ovate, (10–)15–25(–35) × 4–17 mm, margins entire, plane, apex acuminate, surfaces glabrous or adaxial minutely glandular-puberulent.

usually cauline, relatively even-sized or largest proximally and gradually reduced distally;

petiole absent;

blade obovate to narrowly elliptic, (4–)8–54(–70) × (2–)3–23 mm, margins entire or serrate, plane, apex obtuse to rounded, surfaces: proximals glabrous abaxially, distals glandular-pubescent.

Pedicels

1–1.5 mm in fruit.

1–4(–5) mm in fruit.

Flowers

2 per node, or 1 or 2 per node on 1 plant, chasmogamous.

1 per node, chasmogamous.

Styles

pubescent, at least on distal 1/2.

glandular-puberulent.

Corollas

magenta or rose purple, tube yellow, throat yellow, throat and distal tube red-spotted on floor, palate ridges yellow, tube-throat 13–16(–19) mm, limb 16–24 mm diam., bilabiate.

lavender to magenta with diffuse dark markings on sides of darker tube-throat and with dark red-purple midveins on lobes extending from throat, lobes not dark at base, floor white or yellow, fading to white at mouth, palate ridges white or yellow fading to white distally, throat ceiling glabrous, tube-throat 10–20 mm, limb 8–20 mm diam., not bilabiate.

Calyces

symmetrically attached to pedicels, not inflated in fruit, 7–10 mm, glabrous or minutely stipitate-glandular, lobes unequal, apex linear-acuminate, sharp-pointed, ribs green distally, intercostal areas whitish.

symmetrically attached to pedicels, inflated in fruit, (7–)8–15 mm, villous, hairs eglandular, lobes subequal, apex acute to attenuate, ribs and intercostal areas often reddish.

Capsules

10–15 mm.

7–11 mm.

Anthers

included, glabrous or sparsely hirsutulous.

included, ciliate.

Stigmas

exserted, lobes subequal.

included, lobes unequal, abaxial 1.5 times adaxial.

2n

= 16.

Diplacus cusickii

Diplacus viscidus

Phenology Flowering May–Jul. Flowering Apr–Jul.
Habitat Slopes, canyons, washes, ditches, sand talus, diatomaceous slopes, basalt outcrops, black volcanic gravel, volcanic ash and sand, sagebrush areas. Chaparral clearings and openings.
Elevation 800–1000 m. (2600–3300 ft.) 90–1300 m. (300–4300 ft.)
Distribution
from FNA
ID; OR
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
Discussion

Diplacus cusickii is endemic to northern Malheur County, Oregon, and along the Snake River in Ada and Owyhee counties, Idaho. Its narrow geographic range reflects the segregation of the more widely distributed D. cusickioides.

D. M. Thompson (2005) noted that collections in northern Malheur County, Oregon, were intermediate between Mimulus cusickii and M. nanus; the type of M. cusickii is from this area and is among the narrowly endemic, supposedly putative intermediates (G. L. Nesom 2013c). These plants have abruptly and sharply acuminate leaf apices like the more widespread form traditionally identified as Diplacus cusickii; they differ in having glabrous leaf surfaces. Typical D. nanus occurs in close sympatry, without intergradation, with the populations in northern Malheur County. Because of their distinctive morphology and coherent geography, the northern Malheur County plants are reasonably recognized as a distinct species. The more widely distributed form formerly identified as D. cusickii now is identified as D. cusickioides. Populations of D. cusickii in the narrow sense along the Snake River in Ada and Owyhee counties, Idaho, may have dispersed there from the Oregon center. Some plants of D. cusickioides in the Leslie Gulch area of east-central Malheur County have somewhat reduced vestiture, approaching that of D. cusickii.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

D. M. Thompson (2005) noted that Diplacus viscidus and D. compactus (as Mimulus viscidus var. compactus) are parapatric and may intergrade in central Mariposa County. The two taxa are distinguished by the presence or absence of dark stripes on the corolla lobe midveins, which are evident even on herbarium specimens. Thompson found that the two remained distinct when grown together in the greenhouse.

Diplacus viscidus is known from Amador, Calaveras, Eldorado, Mariposa, Merced, and Tuolumne counties; D. compactus continues south through Fresno, northern Kern, Madera, Mariposa, and Tulare counties.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 439. FNA vol. 17, p. 432.
Parent taxa Phrymaceae > Diplacus Phrymaceae > Diplacus
Sibling taxa
D. angustatus, D. aridus, D. aurantiacus, D. bicolor, D. bigelovii, D. bolanderi, D. brandegeei, D. brevipes, D. calycinus, D. cascadensis, D. clevelandii, D. clivicola, D. compactus, D. congdonii, D. constrictus, D. cusickioides, D. deschutesensis, D. douglasii, D. fremontii, D. grandiflorus, D. graniticola, D. jepsonii, D. johnstonii, D. kelloggii, D. layneae, D. leptaleus, D. linearis, D. longiflorus, D. mephiticus, D. mohavensis, D. nanus, D. ovatus, D. parryi, D. parviflorus, D. pictus, D. pulchellus, D. puniceus, D. pygmaeus, D. rattanii, D. rupicola, D. rutilus, D. thompsonii, D. torreyi, D. traskiae, D. tricolor, D. vandenbergensis, D. viscidus
D. angustatus, D. aridus, D. aurantiacus, D. bicolor, D. bigelovii, D. bolanderi, D. brandegeei, D. brevipes, D. calycinus, D. cascadensis, D. clevelandii, D. clivicola, D. compactus, D. congdonii, D. constrictus, D. cusickii, D. cusickioides, D. deschutesensis, D. douglasii, D. fremontii, D. grandiflorus, D. graniticola, D. jepsonii, D. johnstonii, D. kelloggii, D. layneae, D. leptaleus, D. linearis, D. longiflorus, D. mephiticus, D. mohavensis, D. nanus, D. ovatus, D. parryi, D. parviflorus, D. pictus, D. pulchellus, D. puniceus, D. pygmaeus, D. rattanii, D. rupicola, D. rutilus, D. thompsonii, D. torreyi, D. traskiae, D. tricolor, D. vandenbergensis
Synonyms Eunanus cusickii, Mimulus cusickii Mimulus viscidus, M. fremontii var. viscidus, M. subsecundus var. viscidus
Name authority (Greene) G. L. Nesom: Phytoneuron 2012-39: 28. (2012) (Congdon) G. L. Nesom: Phytoneuron 2012-39: 29. (2012)
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