Diplacus cusickii |
Diplacus nanus |
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Cusick's monkey-flower |
dwarf monkey flower, dwarf purple monkey-flower, purple monkeyflower |
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Habit | Herbs, annual. | Herbs, annual. |
Stems | erect to erect-ascending, 10–80 mm, distal internodes 2–20 mm, minutely glandular-puberulent. |
erect, 30–120 mm, minutely glandular-puberulent. |
Leaves | basal and cauline or usually cauline, relatively even-sized or gradually larger distally; petiole absent, proximal base short petiole-like, 1–5 mm; blade ovate to broadly elliptic-ovate, (10–)15–25(–35) × 4–17 mm, margins entire, plane, apex acuminate, surfaces glabrous or adaxial minutely glandular-puberulent. |
basal and cauline, relatively even-sized; petiole absent; blade narrowly elliptic to oblanceolate, ovate, obovate, or elliptic-lanceolate, (1–)3–30(–50) × (0.4–)0.7–8(–20) mm, margins entire, plane, apex rounded or obtuse, surfaces minutely glandular-puberulent. |
Pedicels | 1–1.5 mm in fruit. |
1–3 mm in fruit. |
Flowers | 2 per node, or 1 or 2 per node on 1 plant, chasmogamous. |
2 per node, or 1 or 2 per node on 1 plant, chasmogamous. |
Styles | pubescent, at least on distal 1/2. |
glandular-puberulent or glandular-pubescent. |
Corollas | magenta or rose purple, tube yellow, throat yellow, throat and distal tube red-spotted on floor, palate ridges yellow, tube-throat 13–16(–19) mm, limb 16–24 mm diam., bilabiate. |
magenta to purplish, dark line often extending onto each abaxial lip lobe from throat, palate ridges yellow with red-purple speckling and border, throat floor villous with hairs extending onto abaxial lip, tube 1.1–1.9 mm diam. at filament insertion, tube-throat 11–15 mm, limb 8–14 mm diam., usually, rarely not, bilabiate. |
Calyces | symmetrically attached to pedicels, not inflated in fruit, 7–10 mm, glabrous or minutely stipitate-glandular, lobes unequal, apex linear-acuminate, sharp-pointed, ribs green distally, intercostal areas whitish. |
symmetrically attached to pedicels, not inflated in fruit, 6–9 mm, minutely glandular-puberulent, lobes subequal, apex acute-apiculate, acuminate, or attenuate, ribs dark green or reddish, intercostal areas whitish. |
Capsules | 10–15 mm. |
8–12 mm. |
Anthers | included, glabrous or sparsely hirsutulous. |
included or exserted, ciliate. |
Stigmas | exserted, lobes subequal. |
exserted, lobes equal. |
2n | = 16. |
|
Diplacus cusickii |
Diplacus nanus |
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Phenology | Flowering May–Jul. | Flowering (Apr–)May–Jul. |
Habitat | Slopes, canyons, washes, ditches, sand talus, diatomaceous slopes, basalt outcrops, black volcanic gravel, volcanic ash and sand, sagebrush areas. | Openings in sagebrush, disturbed slopes, granite outcrops. |
Elevation | 800–1000 m. (2600–3300 ft.) | (300–)1100–2300(–2900) m. ((1000–)3600–7500(–9500) ft.) |
Distribution |
ID; OR
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CA; ID; MT; NV; OR; WA; WY
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Discussion | Diplacus cusickii is endemic to northern Malheur County, Oregon, and along the Snake River in Ada and Owyhee counties, Idaho. Its narrow geographic range reflects the segregation of the more widely distributed D. cusickioides. D. M. Thompson (2005) noted that collections in northern Malheur County, Oregon, were intermediate between Mimulus cusickii and M. nanus; the type of M. cusickii is from this area and is among the narrowly endemic, supposedly putative intermediates (G. L. Nesom 2013c). These plants have abruptly and sharply acuminate leaf apices like the more widespread form traditionally identified as Diplacus cusickii; they differ in having glabrous leaf surfaces. Typical D. nanus occurs in close sympatry, without intergradation, with the populations in northern Malheur County. Because of their distinctive morphology and coherent geography, the northern Malheur County plants are reasonably recognized as a distinct species. The more widely distributed form formerly identified as D. cusickii now is identified as D. cusickioides. Populations of D. cusickii in the narrow sense along the Snake River in Ada and Owyhee counties, Idaho, may have dispersed there from the Oregon center. Some plants of D. cusickioides in the Leslie Gulch area of east-central Malheur County have somewhat reduced vestiture, approaching that of D. cusickii. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Diplacus nanus is broadly distributed through northern California, southern Idaho, and eastern Oregon, with stations in Ravalli County, Montana, and Park County, Wyoming, and scattered localities in Washington. Diplacus nanus is generally recognized by its strongly bilabiate corollas with purplish (not yellow) tubes and two dark purple patches along the sides of the throats. The glandular-puberulent vestiture of D. nanus contrasts with the glandular-pubescent and viscid-villous vestiture (with hairs much longer) of D. mephiticus. W. L. Ezell (1971) noted that in the Siskiyou Mountains of Josephine County, Oregon, and adjacent Siskiyou and Trinity counties, California, corollas of Diplacus nanus do not have clearly differentiated abaxial and adaxial lips. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 439. | FNA vol. 17, p. 440. |
Parent taxa | Phrymaceae > Diplacus | Phrymaceae > Diplacus |
Sibling taxa | ||
Synonyms | Eunanus cusickii, Mimulus cusickii | Mimulus nanus, M. tolmiei |
Name authority | (Greene) G. L. Nesom: Phytoneuron 2012-39: 28. (2012) | (Hooker & Arnott) G. L. Nesom: Phytoneuron 2012-39: 29. (2012) |
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